The maximum number of cheliceralcheliceral:
of or pertaining to the chelicera.
segments in mites is three: a basal articlebasal article:
the most basal of the maximum of three segments of the chelicera; usually absent or obscure in Acariformes.
, a middle article that usually ends in a fixed digitfixed digit:
the distal extension of the middle article of the chelicera; usually bearing teeth and a distal hook and opposed to the movable digit in chelate-dentate forms, but often regressed; in Mesostigmata the fixed digit may bear the pilus dentilis.
, and a movable digit that opposes the fixed digitfixed digit:
the distal extension of the middle article of the chelicera; usually bearing teeth and a distal hook and opposed to the movable digit in chelate-dentate forms, but often regressed; in Mesostigmata the fixed digit may bear the pilus dentilis.. However, in some uropodid mites (Fig. 1), the chelicerae are very elongate and the middle article is subdivided.
Most parasitiform mites have 3-segmented chelicerae (Fig. 2), although in ticks (Fig. 3) and some parasitic Mesostigmata, the chelicerae are so modified for blood-feeding that segmentationsegmentation:
in mites distinct external segments have been lost but remnants of segmentation may be represented by hysterosomal folds or transverse arrays of setae and other cuticular sense organs. In theory, all chelicerates have a prosoma composed of 6 segments (cheliceral, pedipalpal, and four leg-bearing segments = body segments I-VI). Ventrally the positions of the prosomal segments can be identified by the insertions of their appendages, but dorsally they are obscured. The opisthosoma is thought to comprise an additional 12-13 segments (body segments VII-XVIII or XIX), but appears to be somewhat to much reduced in most mites, except possibly Opilioacarida. In early derivative Acariformes (e.g., many Endeostigmata), hysterosomal folds are thought to represent segmentation and in the Grandjean system are designated (from the sejugal furrow to the anus): C, D, E, F, H, PS AD, AN, PA. There is disagreement in the literature over the origin of 'segments' C and D. Adherents of Grandjean consider them to be opisthosomatic (with C probably representing a fusion of the pregenital [body segment VII] and genital [VIII] segments). Others believe that C and D are the dorsal regions of the last two prosomal segments that bear leggs III and IV (i.e., body segments V & VI).
is obscured.
Most acariform mites have lost the basalbasal:
towards the base of a structure; on a limb, towards the insertion on the body.
segment and the chelicerae appear to be 2-segmented (Fig. 1), although the regression or loss of the fixed digitfixed digit:
the distal extension of the middle article of the chelicera; usually bearing teeth and a distal hook and opposed to the movable digit in chelate-dentate forms, but often regressed; in Mesostigmata the fixed digit may bear the pilus dentilis. may obscure this and in spider mites (Fig. 4) and their relatives the chelicerae are so modified for feeding on plant cells that segmentationsegmentation:
in mites distinct external segments have been lost but remnants of segmentation may be represented by hysterosomal folds or transverse arrays of setae and other cuticular sense organs. In theory, all chelicerates have a prosoma composed of 6 segments (cheliceral, pedipalpal, and four leg-bearing segments = body segments I-VI). Ventrally the positions of the prosomal segments can be identified by the insertions of their appendages, but dorsally they are obscured. The opisthosoma is thought to comprise an additional 12-13 segments (body segments VII-XVIII or XIX), but appears to be somewhat to much reduced in most mites, except possibly Opilioacarida. In early derivative Acariformes (e.g., many Endeostigmata), hysterosomal folds are thought to represent segmentation and in the Grandjean system are designated (from the sejugal furrow to the anus): C, D, E, F, H, PS AD, AN, PA. There is disagreement in the literature over the origin of 'segments' C and D. Adherents of Grandjean consider them to be opisthosomatic (with C probably representing a fusion of the pregenital [body segment VII] and genital [VIII] segments). Others believe that C and D are the dorsal regions of the last two prosomal segments that bear leggs III and IV (i.e., body segments V & VI).
is obscured. In heterostigmatine Prostigmata, the chelicerae are hidden within a gnathosomal capsulegnathosomal capsule:
a fusion of the gnathosomal elements (chelicerae and subcapitulum) into a single structure, as seen in protigmatans such as Heterostigmata, Myobiidae, and Cheyletoidea (also tegmen).
(Fig. 5), but in the Astigmata heteromorphicheteromorphic:
having different morphological forms; referring either to different forms within a particular life stage (e.g., normal and heteromorphic deutonymphs in some Mesostigmata; protogynes vs. deutogynes in Eriophyoidea; heteromorphic vs. homeomorphic males in the Astigmata) or to a developmental stage that differs radically from other stages (e.g., the heteromorphic deutonymph or hypopus in the Astigmata).
stages the chelicerae are truly absent.
Author: David E. Walter
Content last updated Sept. 2006, unless indicated otherwise
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