KEYS    About TortAI    Fact Sheets    Glossary    ID Thumbnails    DNA Search    Dissection Guides Archipini << Previous Next >> CAPS Primary Target - Adult CAPS Secondary Target - Adult CAPS Non-target - Adult Port Interception Target - Larva Fig. 1: Epichoristodes acerbella Fig. 1: Epichoristodes acerbella Fig. 2: Archips argyrospila Fig. 2: Archips argyrospila Fig. 3: Pandemis cerasana Fig. 3: Pandemis cerasana Fig. 4: Argyrotaenia franciscana Fig. 4: Argyrotaenia franciscana Fig. 5: Epiphyas postvittana Fig. 5: Epiphyas postvittana Fig. 6: Isotenes miserana Fig. 6: Isotenes miserana Fig. 7: Ctenopseustis obliquana Fig. 7: Ctenopseustis obliquana Fig. 8: Adoxophyes orana Fig. 8: Adoxophyes orana Fig. 9: Clepsis peritana Fig. 9: Clepsis peritana Fig. 10: Archips podana Fig. 10: Archips podana Fig. 11: Cacoecimorpha pronubana Fig. 11: Cacoecimorpha pronubana Fig. 12: Choristoneura rosaceana Fig. 12: Choristoneura rosaceana Overview The Archipini is the largest tribe within the Tortricinae, with more than 1,600 described species in 150 genera. Its members are present in all regions but are uncommon in the neotropics. Forewing pattern varies greatly; however, a large number of species have a generalized fasciate pattern, with a well-expressed basal and median fascia, and a dark costal spot. Male genitalia are characterized by an elongate, variably-shaped uncus, well-developed gnathos, and rounded, sometimes membranous or folded, valvae. Females of nearly all Archipini have a daggar-shaped signum with a capitulum; this character is the only true synapomorphy for the group. This group contains the most pest species of any tortricid tribe. Larvae are mostly polyphagous, external feeders, and some species, such as the light brown apple moth (Epiphyas postvittana), have been recorded feeding on more than 500 species of plants. Larvae of many species have few diagnostic morphological characters, and it may be difficult to separate larvae of various genera such as Archips, Argyrotaenia, Clepsis, etc. As such, many larval interceptions at U.S. ports of entry are not identified below the subfamily level except in certain cases of specific host/origin associations. Nearly half of the taxa treated on this site are archipines. These taxa include a mix of target and non-target species; they are arranged below by genus. For photos of additional non-targets not covered here, visit the Moth Photographers Group link below. Adoxophyes Adoxophyes spp Adoxophyes furcatana Adoxophyes honmai Adoxophyes negundana Adoxophyes orana Archips Archips spp Archips argyrospila Archips cerasivorana Archips crataegana Archips fuscocupreanus Archips grisea Archips mortuana Archips podana Archips purpurana Archips rosana Archips semiferanus Archips xylosteana Argyrotaenia Argyrotaenia spp Argyrotaenia franciscana Argyrotaenia ljungiana Argyrotaenia mariana Argyrotaenia quadrifasciana Argyrotaenia quercifoliana Argyrotaenia velutinana Cacoecimorpha Cacoecimorpha pronubana Choristoneura Choristoneura occidentalis Choristoneura rosaceana Clepsis Clepsis spp Clepsis clemensiana Clepsis fucana Clepsis melaleucanus Clepsis peritana Clepsis persicana Clepsis spectrana Clepsis virescana Ctenopseustis Ctenopseustis obliquana Epichoristodes Epichoristodes acerbella Epiphyas Epiphyas postvittana Isotenes Isotenes miserana Pandemis Pandemis cerasana Pandemis heparana Pandemis lamprosana Pandemis limitata Pandemis pyrusana Planotortrix Planotortrix excessana Xenotemna Xenotemna pallorana Links The following external links require an internet connection. The authors, CSU, CDFA, and USDA/APHIS/PPQ/CPHST are not responsible for the availability or content of these external sites: Moth Photographers Group - Archipini References Dombroskie, J. J. 2011. Aspects of archipine evolution (Lepidoptera: Tortricidae). University of Alberta, Department of Biological Sciences. Ph.D. dissertation. 488 pp. Freeman, T. N. 1958. The Archipinae of North America (Lepidoptera: Tortricidae). The Canadian Entomologist Supplement 7 (Vol. 90):  1-89. Horak, M. and R. L. Brown. 1991. Taxonomy and phylogeny, pp. 23-50. In: L. P. S. van der Geest, H. H. Evenhuis (eds.), Tortricid Pests: Their Biology, Natural Enemies and Control. Elsevier, Amsterdam, The Netherlands. Razowski, J. 1987. The genera of Tortricidae (Lepidoptera). Part 1. Palearctic Chlidanotinae and Tortricinae. Acta Zoologica Cracoviensia. 30: 141-355. Tortricids of Agricultural Importance by Todd M. Gilligan and Marc E. Epstein Interactive Keys developed in Lucid 3.5. Last updated August 2014.