Gamasodes

Mesostigmata

t

Taxonomy

updated 2025

Name

Gamasodes Oudemans

Classification

Superorder Parasitiformes » Order Mesostigmata » Suborder Monogynaspida » Infraorder Gamasina » Hyporder Parasitiae » Superfamily Parasitoidea » Family Parasitidae » Subfamily Parasitinae » Genus Gamasodes

Diagnostic characters

Female schizodorsal or divided dorsal shielddivided dorsal shield:
in Mesostigmata, adult mites have two usually subequal dorsal shields.
with setae of dorsal hexagondorsal hexagon:
a fancied hexagonal area staked out by setal pairs j5z5 and j6 in the podonotal region of many mesostigmatans; useful for orienting to setae to determine designations (see Lindquist-Evans system);  setae j3-5 may also form a hexagonal array.
(j5, z5, j6) dissimilar in form; opisthogastric shieldopisthogastric shield:
in Mesostigmata, a shield that covers all of the venter behind legs IV including the anal region; a ventrianal shield that incorporates the metapodal and usually the exopodal elements (e.g., in Parasitidae).
with fewer than 18 pairs of setae, genital region Λ-shaped, with a single anterior point; setae al1, al2 on palppalp:
(= pedipalp) the second pair of limbs in arachnids, used in feeding and originating on either side of the chelicerae.  In mites, the palps may be vestigial, with only a few segments, or have a maximum of  5 freely articulating segments (rarely the femur is subdivided) and a distal or subdistal apotele.
genu in deutonymphs and adults spatulatespatulate:
(Latin spatula = spoon) spatula-shaped; variously interpreted, but usually referring to setae that are flattened and somewhat expanded distally.
; corniculicorniculus:
(pl. corniculi) (also, external malae) a usually horn-like process (sometimes toothed, bifurcate, trifurcate, spine-like, spatulate, or membranous) on the subcapitulum of parasitiform mites that usually supports the salivary styli.  A toothed corniculus could be confused with a rutellum, a possibly homologous structure in Acariformes and Opilioacarida. Phytoseiid image is spatulate corniculus.
short, not extending to anterior margin of palppalp:
(= pedipalp) the second pair of limbs in arachnids, used in feeding and originating on either side of the chelicerae.  In mites, the palps may be vestigial, with only a few segments, or have a maximum of  5 freely articulating segments (rarely the femur is subdivided) and a distal or subdistal apotele.
femur; tritosternumtritosternum:
the sternum of the 3rd body segment (between legs I); produced as a biflagellate structure in Mesostigmata, although sometimes the flagellae (laciniae) are partially or completely fused.  
of male absent or rudimentary; deutonymphdeutonymph:
(also deuteronymph) the second nymphal stage or instar.
(and sometimes female) with spurs on leg II femurfemur:
(pl. femora) major leg segment between trochanter and genu; often subdivided into a basifemur and a telofemur.
(on tubercles), genugenu:
(pl. genua) (= patella) the 4th leg segment, between the femur and tibia.
, and usually tibiatibia:
(pl. tibiae) the leg segment between the genu and the tarsus.
and tarsustarsus:
(pl. tarsi) the subdistal leg segment between the tibia and the pretarsus (apotele).
.

Diagnostic characters of deutonymphs

  • Dorsaldorsal:
    relating to the upper or back side; opposed to ventral.
    shield divided
  • Seta z5 simplesimple:
    unadorned; simple setae are needle-like and without hairs or pectins.
    or stout and distally pilosepilose:
    with a covering of hairs or hair-like processes.
  • Spurs on leg II femurfemur:
    (pl. femora) major leg segment between trochanter and genu; often subdivided into a basifemur and a telofemur.
    (on tubercles), genugenu:
    (pl. genua) (= patella) the 4th leg segment, between the femur and tibia.
    , and usually tibiatibia:
    (pl. tibiae) the leg segment between the genu and the tarsus.
    and tarsustarsus:
    (pl. tarsi) the subdistal leg segment between the tibia and the pretarsus (apotele).

Key to some Gamasodes deutonymphs

  1. Tibiatibia:
    (pl. tibiae) the leg segment between the genu and the tarsus.
    II with 1 ventralventral:
    relating to the lower or under side; opposed to dorsal.
    spur; > 2 pairs of dorsaldorsal:
    relating to the upper or back side; opposed to ventral.
    setae stout and distally pilosepilose:
    with a covering of hairs or hair-like processes.
    ............. 2
    Tibiatibia:
    (pl. tibiae) the leg segment between the genu and the tarsus.
    II with 2 ventralventral:
    relating to the lower or under side; opposed to dorsal.
    spurs; only dorsaldorsal:
    relating to the upper or back side; opposed to ventral.
    setae j1 and r3 stout and distally pilosepilose:
    with a covering of hairs or hair-like processes.
    ............. G. bispinosus (Halbert)

  2. With more than one spurspur:
    a projection, usually tooth- or spine-like, from the body or limbs.
    on femurfemur:
    (pl. femora) major leg segment between trochanter and genu; often subdivided into a basifemur and a telofemur.
    II ............. Gamasodes spp.
    -   With a spine spurspur:
    a projection, usually tooth- or spine-like, from the body or limbs.
    on the venterventer:
    the lower or under side; opposed to dorsum.
    of femurfemur:
    (pl. femora) major leg segment between trochanter and genu; often subdivided into a basifemur and a telofemur.
    II............. 3

  3. Spurspur:
    a projection, usually tooth- or spine-like, from the body or limbs.
    on femurfemur:
    (pl. femora) major leg segment between trochanter and genu; often subdivided into a basifemur and a telofemur.
    II straight; tectumgnathotectum:
    (= epistome, tectum) an often membranous dorsal projection of the anterior margin of the basis capitulum in Mesostigmata; often diagnostic at family-, genus- or species-levels; also often difficult to see without high magnification and good optics.
    with median process and weakly lateral teeth............. G. fimbriatus Karg
    Spurspur:
    a projection, usually tooth- or spine-like, from the body or limbs.
    on femurfemur:
    (pl. femora) major leg segment between trochanter and genu; often subdivided into a basifemur and a telofemur.
    II curved; tectumgnathotectum:
    (= epistome, tectum) an often membranous dorsal projection of the anterior margin of the basis capitulum in Mesostigmata; often diagnostic at family-, genus- or species-levels; also often difficult to see without high magnification and good optics.
    clearly tripartite............. G. spiniger (Trägårdh)

Similar taxa

Key to some Parasitine genera (adult females) (modified from Hyatt 1980)

  1. Lateral setae of palppalp:
    (= pedipalp) the second pair of limbs in arachnids, used in feeding and originating on either side of the chelicerae.  In mites, the palps may be vestigial, with only a few segments, or have a maximum of  5 freely articulating segments (rarely the femur is subdivided) and a distal or subdistal apotele.
    genu (al12) entireentire:
    a shield or sclerite with a continuous margin without incisions.
    , either setiform or spatulatespatulate:
    (Latin spatula = spoon) spatula-shaped; variously interpreted, but usually referring to setae that are flattened and somewhat expanded distally.
    .............................. 2
    - Lateral setae of palppalp:
    (= pedipalp) the second pair of limbs in arachnids, used in feeding and originating on either side of the chelicerae.  In mites, the palps may be vestigial, with only a few segments, or have a maximum of  5 freely articulating segments (rarely the femur is subdivided) and a distal or subdistal apotele.
    genu (al12) bifid.............................. Eugamasus

  2. Posterior ventral regionventral region:
    in Mesostigmata, the area between the genital and anal regions.
    (opisthogaster) with 30 or fewer pairs of setae; associated with a variety of insects, but usually not bumblebees (Bombus).............................. 3
    - Opisthogaster with extensive hypertrichy (>40 pairs of setae); associated with bumblebees (Bombus) or their nests.............................. Parasitellus

  3. Corniculicorniculus:
    (pl. corniculi) (also, external malae) a usually horn-like process (sometimes toothed, bifurcate, trifurcate, spine-like, spatulate, or membranous) on the subcapitulum of parasitiform mites that usually supports the salivary styli.  A toothed corniculus could be confused with a rutellum, a possibly homologous structure in Acariformes and Opilioacarida. Phytoseiid image is spatulate corniculus.
    of normal length, not reaching the anterior margin of the palppalp:
    (= pedipalp) the second pair of limbs in arachnids, used in feeding and originating on either side of the chelicerae.  In mites, the palps may be vestigial, with only a few segments, or have a maximum of  5 freely articulating segments (rarely the femur is subdivided) and a distal or subdistal apotele.
    femur; anterior margin of anterior dorsaldorsal:
    relating to the upper or back side; opposed to ventral.
    shield convex.............................. 4
    - Corniculicorniculus:
    (pl. corniculi) (also, external malae) a usually horn-like process (sometimes toothed, bifurcate, trifurcate, spine-like, spatulate, or membranous) on the subcapitulum of parasitiform mites that usually supports the salivary styli.  A toothed corniculus could be confused with a rutellum, a possibly homologous structure in Acariformes and Opilioacarida. Phytoseiid image is spatulate corniculus.
    elongate, extending beyond palppalp:
    (= pedipalp) the second pair of limbs in arachnids, used in feeding and originating on either side of the chelicerae.  In mites, the palps may be vestigial, with only a few segments, or have a maximum of  5 freely articulating segments (rarely the femur is subdivided) and a distal or subdistal apotele.
    femur-genugenu:
    (pl. genua) (= patella) the 4th leg segment, between the femur and tibia.
    articulation; margin of anterior dorsaldorsal:
    relating to the upper or back side; opposed to ventral.
    shield concave.............................. Cornigamasus

  4. Genital shieldgenital shield:
    a shield or shields covering the genital opening; in female mongynaspine Mesostigmata this shield is usually called the epigynal (epigynial) shield.  
    (or genital region of hologastric shieldhologastric shield:
    in Mesostigmata, a shield that covers the genital, ventral and anal regions; a ventrianal shield that incorporates the metapodal elements and is fused to the epigynal shield. 
    ) subtriangularsubtriangular:
    more or less triangular.  
    , acuminate.............................. 5
    - Genital region of hologastric shieldhologastric shield:
    in Mesostigmata, a shield that covers the genital, ventral and anal regions; a ventrianal shield that incorporates the metapodal elements and is fused to the epigynal shield. 
    distally tricuspidtricuspid:
    having three points or cusps, e.g., the gnathotectum of some Mesostigmata.
    .............................. Porrhostaspis

  5. Lateral seta of palppalp:
    (= pedipalp) the second pair of limbs in arachnids, used in feeding and originating on either side of the chelicerae.  In mites, the palps may be vestigial, with only a few segments, or have a maximum of  5 freely articulating segments (rarely the femur is subdivided) and a distal or subdistal apotele.
    femur (al) entireentire:
    a shield or sclerite with a continuous margin without incisions.
    , usually spatulatespatulate:
    (Latin spatula = spoon) spatula-shaped; variously interpreted, but usually referring to setae that are flattened and somewhat expanded distally.
    .............................. 8
    - Lateral seta of palppalp:
    (= pedipalp) the second pair of limbs in arachnids, used in feeding and originating on either side of the chelicerae.  In mites, the palps may be vestigial, with only a few segments, or have a maximum of  5 freely articulating segments (rarely the femur is subdivided) and a distal or subdistal apotele.
    femur bifid, serrateserrate:
    with closely set teeth that resemble the cutting edge of a saw.
    , or serrateserrate:
    with closely set teeth that resemble the cutting edge of a saw.
    -spatulate.............................. 6

  6. Setae of dorsal hexagondorsal hexagon:
    a fancied hexagonal area staked out by setal pairs j5z5 and j6 in the podonotal region of many mesostigmatans; useful for orienting to setae to determine designations (see Lindquist-Evans system);  setae j3-5 may also form a hexagonal array.
    dissimilar, seta z5 longer and usually thicker than j5 and j6............................... 7
    - Setae of dorsal hexagondorsal hexagon:
    a fancied hexagonal area staked out by setal pairs j5z5 and j6 in the podonotal region of many mesostigmatans; useful for orienting to setae to determine designations (see Lindquist-Evans system);  setae j3-5 may also form a hexagonal array.
    similar in size and ornamentation.............................. Vulgarogamasus

  7. Sternal shieldsternal shield:
    a shield in the anterior intercoxal region of parasitiform mites that bears one or more pairs of sternal setae.
    entire; seta z5 usually thicker than j5 and j6 and distally pilosepilose:
    with a covering of hairs or hair-like processes.
    associated with dung (and dung beetles), compost, etc.............................. Parasitus
    - Sternal shieldsternal shield:
    a shield in the anterior intercoxal region of parasitiform mites that bears one or more pairs of sternal setae.
    split medially; seta z5 usually longer than j5 and j6 but similar in form; associated with bark beetles.............................. Schizosthetus

  8. Dorsaldorsal:
    relating to the upper or back side; opposed to ventral.
    shield completely divided.............................. 9
    - Schizodorsal shieldschizodorsal shield:
    in Mesostigmata, a holodorsal shield that incised laterally at about the midpoint or podonotal and opisthonotal shields that are fused only medially.
    .............................. Gamasodes bispinosus group

  9. Seta z5 of dorsal hexagondorsal hexagon:
    a fancied hexagonal area staked out by setal pairs j5z5 and j6 in the podonotal region of many mesostigmatans; useful for orienting to setae to determine designations (see Lindquist-Evans system);  setae j3-5 may also form a hexagonal array.
    thickened and distally pilosepilose:
    with a covering of hairs or hair-like processes.
    ; associated with various insects.............................. 10
    - Seta z5 simplesimple:
    unadorned; simple setae are needle-like and without hairs or pectins.
    ; associated with carrion beetles (Silphidae).............................. Poecilochirus

  10. Articulationarticulation:
    a region of differentiated cuticle joining two parts of an exoskeleton; a joint, scissure or furrow.
    between sternal and metasternal shields oblique; ambulacra of legs II–IV simplesimple:
    unadorned; simple setae are needle-like and without hairs or pectins.
    , rounded.............................. Gamasodes
    - Articulationarticulation:
    a region of differentiated cuticle joining two parts of an exoskeleton; a joint, scissure or furrow.
    (or suturesuture:
    an inflexible juncture between two body parts; line of fusion between two formerly separate body parts or regions.  This term is often misused for flexible articulations.
    ) between sternal and metasternal shields transverse; ambulacra of legs II–IV long and acuminate.............................. Trachygamasus

Ecology and distribution

Soil-inhabiting predators of small arthropods and other invertebrates; several species have been found in the nests of birds and mammals and one species in rotting fungal sporocarps; phoretic on muscid and drosophilid flies (rarely on dung beetles).

Known species

ca. 25 described species: Gamasodes aequipilis Athias-Henriot, 1980 (Iceland), Gamasodes assamensis Bhattacharyya, 1971Bhattacharyya, 1971:
Bhattacharyya SK. 1971. Studies on Indian mites (Acarina: Mesostigmata): 10. One new species and five records from Assam. Oriental Insects 5: 495-500.
(India), Gamasodes bispinosus (Halbert, 1915Halbert, 1915:
Halbert JN. 1915. Claire Island Survey. Acarinida Section II - Terrestrial and marine Acarina. Proceedings of the Royal Irish Academy 31 (39): 45-136.
) (Europe, USSR), Gamasodes buettikeri Samšiňák, 1979Samšiňák, 1979:
Samšiňák K. 1979. Mites of Saudi Arabia Acari from Musca domestica. Fauna Saudi Arabia 1: 69-74.
(Saudi Arabia), Gamasodes bulgatus Athias-Henriot, 1979Athias-Henriot, 1979:
Athias-Henriot C. 1979. Sur le groupe d’espèces “bispinosus” du genre Gamasodes Oudemans, 1939 (Parasitiformes, Parasitidae). Bulletin Société Zoologique du France 104: 209-220.
(Canary Islands), Gamasodes coprophilus Chelebiev, 1980Chelebiev, 1980:
Chelebiev KA. 1980. On mites of the genus Gamasodes (Parasitiformes, Parasitidae) from central Kazakhstan. Zoologicheskii Zhurnal 59: 1098-1100.
(USSR), Gamasodes corniculans Athias-Henriot, 1978Athias-Henriot, 1978:
Athias-Henriot C. 1978. Deutonymphen der Gattung Gamasodes Oudms (Parasitiformes, Parasitidae) aus der Mongolei. Annales Historico-Naturales Musei Nationalis Hungarici 70: 345-348.
(Mongolia), Gamasodes diceras Athias-Henriot, 1980 (Europe), Gamasodes fimbriatus Karg, 1971Karg, 1971:
Karg W. 1971. Acari (Acarina), Milben Unterordnung Anactinochaeta (Parasitiformes) Die freilebenden Gamasina (Gamasides), Raubmilben. Die Tierwelt Deutschlands 59: 1-475.
(Europe), Gamasodes guoluoensis Gu and Liu, 1995Gu and Liu, 1995:
Gu Y-M and Liu J-O. 1995. A new species of the genus Gamasodes from China (Acari: Parasitidae). Acta Zootaxonomica Sinica 20: 65-67.
(China), Gamasodes hortivagus (Berlese, 1903Berlese, 1903:
Berlese A. 1903. Acari nuovi Manipulus I us. Redia 1: 235-252.
) (Europe, Algeria, Canary Islands [= Gamasodes cabylus Athias-Henriot 1961], Gamasodes ignoratus Oudemans, 1939bOudemans, 1939b:
Oudemans AC. 1939b. Neue funde auf dem gebiete der systematik und der nomenklatur der Acari IV. Zoologischer Anzeiger 126: 195-201.
(Europe), Gamasodes inermis Athias-Henriot, 1978Athias-Henriot, 1978:
Athias-Henriot C. 1978. Deutonymphen der Gattung Gamasodes Oudms (Parasitiformes, Parasitidae) aus der Mongolei. Annales Historico-Naturales Musei Nationalis Hungarici 70: 345-348.
(Mongolia), Gamasodes marmotae Ma, 1992Ma, 1992:
Ma LM. 1992. Descriptions of a new species and an unknown male of Gamasodes from north Qing-Zang Plateau (Acari: Mesostigmata, Parasitidae). Acta Entomologica Sinica 35: 113-116.
(China), Gamasodes micherdzinskii Davydova, 1973Davydova, 1973:
Davydova MS. 1973. Description of new species from the genus Gamasodes and the male of Veigaia beklemischevi (Parasitiformes, Gamasoidea). pp. 103-112. In: Cherepanov AI, ed. New and little-known species of siberian fauna, Vol. 6. Nauka, Novosibirsk.
(USSR, China), Gamasodes miliaris Athias-Henriot, 1980 (Morocco), Gamasodes nudus Tseng, 1995Tseng, 1995:
Tseng YH. 1995. A taxonomical study of free-living gamasine mite family Parasitidae Oudemans (Acari, Mesostigmata) from Taiwan. Journal of the Taiwan Museum 48: 11-81.
(Taiwan), Gamasodes plenigranosus Athias-Henriot, 1979Athias-Henriot, 1979:
Athias-Henriot C. 1979. Sur le groupe d’espèces “bispinosus” du genre Gamasodes Oudemans, 1939 (Parasitiformes, Parasitidae). Bulletin Société Zoologique du France 104: 209-220.
(Madagascar), Gamasodes poppei (Oudemans, 1903Oudemans, 1903:
Oudemans AC. 1903. Acarologische Aanteekeningen. VI. Entomologische Berichten 1 (12): 83-88.
) (Europe), Gamasodes queenslandicus Halliday and Walter, sp. nov., Gamasodes simplex Athias-Henriot, 1978Athias-Henriot, 1978:
Athias-Henriot C. 1978. Deutonymphen der Gattung Gamasodes Oudms (Parasitiformes, Parasitidae) aus der Mongolei. Annales Historico-Naturales Musei Nationalis Hungarici 70: 345-348.
(Mongolia), Gamasodes sinicus Tian and Gu, 1991Tian and Gu, 1991:
Tian QY and Gu YM. 1991. The first record of Gamasodes in China with description of a new species (Acari: Parasitidae). Acta Zootaxonomica Sinica 16: 432-435.
(China), Gamasodes spiniger (Trägårdh 1910Trägårdh 1910:
Trägårdh I. 1910. Acariden aus dem Sarekgebirge. Naturwissenschaftlichen Naturwissenschaftliche Untersuchungen des Sarakgebirges in Schwedisch-Lappland geleitet von Dr Axel Hamberg. Zoology, Stockholm 4 (Lief 4): 375-586.
)
(Europe) type species [ = Gamasoides spiniger Oudemans 1936Oudemans 1936:
Oudemans AC. 1936. Kritisch historisch overzicht der acarologie, Deerde Gedeelte, 1805-1850. Band A (E J Brill, Leiden): 1-430.
(nom nov pro Gamasus spinipes (Koch 1844), Gamasodes berlesei Oudemans 1939bOudemans 1939b:
Oudemans AC. 1939b. Neue funde auf dem gebiete der systematik und der nomenklatur der Acari IV. Zoologischer Anzeiger 126: 195-201.
. nom nov pro Gamasus spinipes (Koch 1844), Parasitus lunarisimilis Schweizer, 1961], Gamasodes tongdensis Li et al., 1999 (China), Gamasodes viretianus Athias-Henriot, 1980 (Europe)

References

  • Athias-Henriot 1961bAthias-Henriot 1961b:
    Athias-Henriot C. 1961b. Physallolaelaps ampulliger Berl et Gamasodes cabylus n. sp. (Parasitiformes: Laelaptidae, Parasitidae). Acarologia 3: 256–264.
  • Athias-Henriot 1978Athias-Henriot 1978:
    Athias-Henriot C. 1978. Deutonymphen der Gattung Gamasodes Oudms (Parasitiformes, Parasitidae) aus der Mongolei. Annales Historico-Naturales Musei Nationalis Hungarici 70: 345–348.
  • Athias-Henriot 1979Athias-Henriot 1979:
    Athias-Henriot C. 1979. Sur le groupe d’espèces “bispinosus” du genre Gamasodes Oudemans, 1939 (Parasitiformes, Parasitidae). Bulletin Société Zoologique du France 104: 209–220.
  • Athias-Henriot 1980bAthias-Henriot 1980b:
    Athias-Henriot C. 1980b. Recapitulatory note on composition and geography of the genus Gamasodes Ouds (Parasitiformes, Parasitidae), with a description of four new species. Biologisch Jaarboek 48: 50–62.
  • Berlese 1903Berlese 1903:
    Berlese A. 1903. Acari nuovi Manipulus I us. Redia 1: 235–252.
  • Bhattacharyya 1971Bhattacharyya 1971:
    Bhattacharyya SK. 1971. Studies on Indian mites (Acarina: Mesostigmata): 10. One new species and five records from Assam. Oriental Insects 5: 495–500.
  • Chelebiev 1980Chelebiev 1980:
    Chelebiev KA. 1980. On mites of the genus Gamasodes (Parasitiformes, Parasitidae) from central Kazakhstan. Zoologicheskii Zhurnal 59: 1098–1100.
  • Davydova 1973Davydova 1973:
    Davydova MS. 1973. Description of new species from the genus Gamasodes and the male of Veigaia beklemischevi (Parasitiformes, Gamasoidea). pp. 103–112. In: Cherepanov AI, ed. New and little-known species of siberian fauna, Vol. 6. Nauka, Novosibirsk.
  • Fenda 1999Fenda 1999:
    Fenda P. 1999. First records of mites (Acarina, Mesostigmata) from Slovakia. Biologia 54 (5): 528.
  • Gu and Liu 1995Gu and Liu 1995:
    Gu Y-M, Liu J-O. 1995. A new species of the genus Gamasodes from China (Acari: Parasitidae). Acta Zootaxonomica Sinica 20: 65–67.
  • Halbert 1915Halbert 1915:
    Halbert JN. 1915. Claire Island Survey. Acarinida Section II - Terrestrial and marine Acarina. Proceedings of the Royal Irish Academy 31 (39): 45–136.
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  • Karg 1971Karg 1971:
    Karg W. 1971. Acari (Acarina), Milben Unterordnung Anactinochaeta (Parasitiformes) Die freilebenden Gamasina (Gamasides), Raubmilben. Die Tierwelt Deutschlands 59: 1–475.
  • Koch 1835–1844Koch 1835–1844:
    Koch CL. 1835–1844. Deutschlands Crustaceen, Myriapoden und Arachniden: Ein beitrag zur Deutschen fauna. Pustet, Regensburg.
  • Li et al. 1999Li et al. 1999:
    Li C, Yang XZ, Chen HJ. 1999. A new species of the genus Gamasodes from China (Acari: Parasitidae). Acta Zootaxonomica Sinica 24: 156–158.
  • Ma 1992Ma 1992:
    Ma LM. 1992. Descriptions of a new species and an unknown male of Gamasodes from north Qing-Zang Plateau (Acari: Mesostigmata, Parasitidae). Acta Entomologica Sinica 35: 113–116.
  • Micherdzinski 1969Micherdzinski 1969:
    Micherdzinski W. 1969. Die Familie Parasitidae Oudemans 1901 (Acarina, Mesostigmata). Zakład Zoologii Systematycznej Polskiej Akademii Nauk, Krakow. 690 pp.
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    Oudemans AC. 1903. Acarologische Aanteekeningen. VI. Entomologische Berichten 1 (12): 83–88.
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    Oudemans AC. 1936. Kritisch historisch overzicht der acarologie, Deerde Gedeelte, 1805-1850. Band A (E J Brill, Leiden): 1–430.
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    Oudemans AC. 1939a. Neue funde auf dem gebiete der systematik und der nomenclatur der Acari III. Zoologischer Anzeiger 126: 20–24.
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    Oudemans AC. 1939b. Neue funde auf dem gebiete der systematik und der nomenklatur der Acari IV. Zoologischer Anzeiger 126: 195–201.
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  Gamasodes  female venter
Gamasodes female venter
  Gamasodes  female chelicera
Gamasodes female chelicera
  Gamasodes  female sterno-genital region
Gamasodes female sterno-genital region
 Gamasodes  deutonymph
Gamasodes deutonymph