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CAPS Non-target - Adult

Archips grisea (Robinson) (Tortricidae: Tortricinae: Archipini)

Common names: black-shield leaf roller

Synonyms: brauniana (Cacoecia)

Fig. 1: Male

Fig. 1: Male

Fig. 2: Female

Fig. 2: Female

Fig. 3: Female

Fig. 3: Female

Fig. 4: Male genitalia

Fig. 4: Male genitalia

Fig. 5: Female genitalia

Fig. 5: Female genitalia

Adult Recognition

FWL: 7.5-9.0 mm (male); 8.0-11.0 mm (female)

Forewing color is primarily brownish gray, with dark-brown to black markings that are more prominently expressed in the male. Hindwings are a uniform gray. Males have a forewing costal fold.

Adult A. grisea are unlikely to be confused with other native Nearctic Archips. General forewing pattern is similar to some forms of Archips podana, Archips crataegana, and Archips rosana, although the grayish appearance of the forewing and hindwing should be sufficient to distinguish A. grisea from these other species.

Larval Morphology

Larvae are 15-23 mm in length with a green abdomen. The prothoracic shield and head are uniformly black in the last instar. The prothoracic legs are brown to black while the other thoracic legs are pale green and unmarked.

The life cycle and larval habits of A. grisea are similar to those of Archips argyrospila, Archips mortuana, and Choristoneura rosaceana. Larvae of A. grisea are distinguished from these other species by a uniformly black prothoracic shield. Archips cerasivorana larvae also have a uniformly black prothoracic shield, but the larvae live and feed in communal nests.


Archips grisea completes one generation per year. Adults are present in June and July. This species has the same basic life history information as Archips argyrospila and can often be found in association with A. argyrospila infestations.

Females lay eggs in masses of approximately 50-60 individual eggs on small diameter branches of the host. Eggs overwinter and first instar larvae hatch the following spring. Larvae feed on leaves and young fruit and pupation occurs within the final larval feeding site.

Host plants

Chapman and Lienk (1971) state that apple and hawthorn are the primary larval hosts in the northeastern U.S. Other host plants are listed here.

Family Genus/species Common name
Asteraceae Rudbeckia L. coneflower
Fagaceae Quercus L. oak
Juglandaceae Carya Nutt. hickory
Rosaceae Crataegus L. hawthorn
Rosaceae Malus Mill. apple
Rosaceae Prunus virginiana L. chokecherry
Rosaceae Pyrus L. pear


Archips grisea is distributed in the eastern United States, from Maine south to Texas. There is a record from Utah and Chapman and Lienk (1971) speculate this might have been the result of an introduction of infested plants from the East.


Chapman, P. J. and S. E. Lienk. 1971. Tortricid fauna of apple in New York (Lepidoptera: Tortricidae); including an account of apple's occurrence in the state, especially as a naturalized plant. Spec. Publ. Geneva, NY: New York State Agricultural Experiment Station. 122 pp.

Freeman, T. N. 1958. The Archipinae of North America (Lepidoptera: Tortricidae). The Canadian Entomologist Supplement 7 (Vol. 90): 1-89.

Tortricids of Agricultural Importance by Todd M. Gilligan and Marc E. Epstein
Interactive Keys developed in Lucid 3.5. Last updated August 2014.