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Drone flying to mate with virgin; photo by The BeeMD photo collection Drone flying to mate with virgin; photo by The BeeMD photo collection
Mating flight
Drone flying to mate with virgin; photo by The BeeMD photo collection
Queen returning form mating flight; photo courtesy Rusty Burlew, https://www.honeybeesuite.com Queen returning form mating flight; photo courtesy Rusty Burlew, https://www.honeybeesuite.com
Mating flight
Queen returning form mating flight; photo courtesy Rusty Burlew, https://www.honeybeesuite.com
Diagram of DCA: queens mate with drones from colonies other than her own; illustration courtesy Carolina Honeybees, IIC Diagram of DCA: queens mate with drones from colonies other than her own; illustration courtesy Carolina Honeybees, IIC
Mating flight
Diagram of DCA: queens mate with drones from colonies other than her own; illustration courtesy Carolina Honeybees, IIC
Evidence of mice gnawing on entrance reducer; photo by Rusty Burlew Evidence of mice gnawing on entrance reducer; photo by Rusty Burlew
Mice
Evidence of mice gnawing on entrance reducer; photo by Rusty Burlew

Mouse in hive: photo by Cathy King Mouse in hive: photo by Cathy King
Mice
Mouse in hive: photo by Cathy King
Mice destroying portion of comb for their nest; photo by Dewey M. Caron Mice destroying portion of comb for their nest; photo by Dewey M. Caron
Mice
Mice destroying portion of comb for their nest; photo by Dewey M. Caron
Mouse at left with nest in hive; photo by Robyn Underwood Mouse at left with nest in hive; photo by Robyn Underwood
Mice
Mouse at left with nest in hive; photo by Robyn Underwood
Evidence of mouse chewing on top bar; photo by Dewey M. Caron Evidence of mouse chewing on top bar; photo by Dewey M. Caron
Mice
Evidence of mouse chewing on top bar; photo by Dewey M. Caron

earwig; photo courtesy Joseph Berger, Bugwood.org earwig; photo courtesy Joseph Berger, Bugwood.org
Miscellaneous animal pests of colonies
earwig; photo courtesy Joseph Berger, Bugwood.org
Cockroach adult, American; photo by Dewey M. Caron Cockroach adult, American; photo by Dewey M. Caron
Miscellaneous animal pests of colonies
Cockroach adult, American; photo by Dewey M. Caron
Scorpion inside top cover of hive; photo by University of Florida Scorpion inside top cover of hive; photo by University of Florida
Miscellaneous animal pests of colonies
Scorpion inside top cover of hive; photo by University of Florida
Skunk visiting colony at night; note digging evidence and scratch marks on colony; photo by University of Delaware Skunk visiting colony at night; note digging evidence and scratch marks on colony; photo by University of Delaware
Miscellaneous animal pests of colonies
Skunk visiting colony at night; note digging evidence and scratch marks on colony; photo by University of Delaware

Woodpecker damage to hive; photo by University of Delaware Woodpecker damage to hive; photo by University of Delaware
Miscellaneous animal pests of colonies
Woodpecker damage to hive; photo by University of Delaware
Spider webbing among un-hatched brood cells from comb in storage; photo by The BeeMD photo collection Spider webbing among un-hatched brood cells from comb in storage; photo by The BeeMD photo collection
Miscellaneous animal pests of colonies
Spider webbing among un-hatched brood cells from comb in storage; photo by The BeeMD photo collection
Moth in hive (probably wax moth); photo by Robert Snyder Moth in hive (probably wax moth); photo by Robert Snyder
Miscellaneous animal pests of colonies
Moth in hive (probably wax moth); photo by Robert Snyder
Fly maggot in bee hive; photo by Dewey M. Caron Fly maggot in bee hive; photo by Dewey M. Caron
Miscellaneous animal pests of colonies
Fly maggot in bee hive; photo by Dewey M. Caron

Crab spider about to grab a forager on Sedum; photo by Kathy Keatley Garvey Crab spider about to grab a forager on Sedum; photo by Kathy Keatley Garvey
Miscellaneous animal pests of colonies
Crab spider about to grab a forager on Sedum; photo by Kathy Keatley Garvey
Larger bodied dragonflies may prey on flying bees; photo by Kathy Keatley Garvey Larger bodied dragonflies may prey on flying bees; photo by Kathy Keatley Garvey
Miscellaneous animal pests of colonies
Larger bodied dragonflies may prey on flying bees; photo by Kathy Keatley Garvey
Moldy bees from winter deadout; photo by Dewey M. Caron Moldy bees from winter deadout; photo by Dewey M. Caron
Mold
Moldy bees from winter deadout; photo by Dewey M. Caron
Moldy bees and comb cells from a winter deadout; photo by Dewey Caron Moldy bees and comb cells from a winter deadout; photo by Dewey Caron
Mold
Moldy bees and comb cells from a winter deadout; photo by Dewey Caron

Mold on deadout frame; photo by Rusty Burlew Mold on deadout frame; photo by Rusty Burlew
Mold
Mold on deadout frame; photo by Rusty Burlew
Excess moisture on bottom board; photo by Dewey M. Caron Excess moisture on bottom board; photo by Dewey M. Caron
Mold
Excess moisture on bottom board; photo by Dewey M. Caron
Moldy dead bees; photo by Dewey M. Caron Moldy dead bees; photo by Dewey M. Caron
Mold
Moldy dead bees; photo by Dewey M. Caron
A two-queen colony; photo by Andrew Abrahams A two-queen colony; photo by Andrew Abrahams
Multiple queens
A two-queen colony; photo by Andrew Abrahams

Nosema spores (1000X); photo by The BeeMD photo collection Nosema spores (1000X); photo by The BeeMD photo collection
Nosema
Nosema spores (1000X); photo by The BeeMD photo collection
Nosema infection of midgut (top), normal midgut (bottom); photo by USDA Nosema infection of midgut (top), normal midgut (bottom); photo by USDA
Nosema
Nosema infection of midgut (top), normal midgut (bottom); photo by USDA
Midgut with Nosema (top); photo by Zachary Huang, beetography.com Midgut with Nosema (top); photo by Zachary Huang, beetography.com
Nosema
Midgut with Nosema (top); photo by Zachary Huang, beetography.com
Feeding bees Nosema spores to study disease; photo by Steve Ausmus, USDA Feeding bees Nosema spores to study disease; photo by Steve Ausmus, USDA
Nosema
Feeding bees Nosema spores to study disease; photo by Steve Ausmus, USDA

Weak spring colony can be a sign of nutritional stress; photo by Dewey M. Caron Weak spring colony can be a sign of nutritional stress; photo by Dewey M. Caron
Nutritional stress
Weak spring colony can be a sign of nutritional stress; photo by Dewey M. Caron
Nutritional stress; too little jelly in larval cells (see arrows); photo by Dewey M. Caron Nutritional stress; too little jelly in larval cells (see arrows); photo by Dewey M. Caron
Nutritional stress
Nutritional stress; too little jelly in larval cells (see arrows); photo by Dewey M. Caron
Larvae (at arrows) attempting to crawl out of cells, likely due to starvation; photo by Oregon State University Larvae (at arrows) attempting to crawl out of cells, likely due to starvation; photo by Oregon State University
Nutritional stress
Larvae (at arrows) attempting to crawl out of cells, likely due to starvation; photo by Oregon State University
Orientation flights in front of a hive with flow super; photo by Dewey M. Caron Orientation flights in front of a hive with flow super; photo by Dewey M. Caron
Orientation flight
Orientation flights in front of a hive with flow super; photo by Dewey M. Caron