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Dead capped brood (see arrows) on frame in spring deadout, likely from cluster moving away from brood; photo by Dewey M. Caron Dead capped brood (see arrows) on frame in spring deadout, likely from cluster moving away from brood; photo by Dewey M. Caron
Chilled brood
Dead capped brood (see arrows) on frame in spring deadout, likely from cluster moving away from brood; photo by Dewey M. Caron
Chilled, capped brood; deadout; photo by Charles Vanden Heuvel Chilled, capped brood; deadout; photo by Charles Vanden Heuvel
Chilled brood
Chilled, capped brood; deadout; photo by Charles Vanden Heuvel
Decomposed black brood cells characteristic of chilled brood; photo by University of Delaware Decomposed black brood cells characteristic of chilled brood; photo by University of Delaware
Chilled brood
Decomposed black brood cells characteristic of chilled brood; photo by University of Delaware
Remains of chilled brood; note black material in the bottom of cells; photo by Rick Olson Remains of chilled brood; note black material in the bottom of cells; photo by Rick Olson
Chilled brood
Remains of chilled brood; note black material in the bottom of cells; photo by Rick Olson

Bees with CBPV; photo courtesy Animal and Plant Health Agency, Crown Copyright, UK Bees with CBPV; photo courtesy Animal and Plant Health Agency, Crown Copyright, UK
Chronic bee paralysis virus
Bees with CBPV; photo courtesy Animal and Plant Health Agency, Crown Copyright, UK
Shiny, greasy adult bees with CBPV; photo Courtesy Animal and Plant Health Agency, Crown Copyright, UK Shiny, greasy adult bees with CBPV; photo Courtesy Animal and Plant Health Agency, Crown Copyright, UK
Chronic bee paralysis virus
Shiny, greasy adult bees with CBPV; photo Courtesy Animal and Plant Health Agency, Crown Copyright, UK
Bee with DWV and cloudy-appearing wings; photo by The BeeMD photo collection Bee with DWV and cloudy-appearing wings; photo by The BeeMD photo collection
Cloudy wing virus
Bee with DWV and cloudy-appearing wings; photo by The BeeMD photo collection
Bee with DWV but also perhaps cloudy wing virus; photo by Robert Snyder Bee with DWV but also perhaps cloudy wing virus; photo by Robert Snyder
Cloudy wing virus
Bee with DWV but also perhaps cloudy wing virus; photo by Robert Snyder

Swarm bivouac cluster; photo by Elaine Timm Swarm bivouac cluster; photo by Elaine Timm
Clustering
Swarm bivouac cluster; photo by Elaine Timm
Small spring cluster: photo by Stephen Repasky Small spring cluster: photo by Stephen Repasky
Clustering
Small spring cluster: photo by Stephen Repasky
Swarm clustered in holly tree; photo by Elaine Timm Swarm clustered in holly tree; photo by Elaine Timm
Clustering
Swarm clustered in holly tree; photo by Elaine Timm
Bees clustering between frames; photo by Lawrence John Connor Bees clustering between frames; photo by Lawrence John Connor
Clustering
Bees clustering between frames; photo by Lawrence John Connor

Remains of a winter cluster; bees died from lack of honey to keep warm; photo by The BeeMD photo collectiono Remains of a winter cluster; bees died from lack of honey to keep warm; photo by The BeeMD photo collectiono
Clustering
Remains of a winter cluster; bees died from lack of honey to keep warm; photo by The BeeMD photo collectiono
View of weak cluster looking upward from bottom bars; photo by Robert Snyder View of weak cluster looking upward from bottom bars; photo by Robert Snyder
Clustering
View of weak cluster looking upward from bottom bars; photo by Robert Snyder
Colony collapse disorder; photo by The BeeMD collection Colony collapse disorder; photo by The BeeMD collection
Colony collapse disorder
Colony collapse disorder; photo by The BeeMD collection
Normal (top), Colony collapse disorder (bottom); photo by 21st Centech Normal (top), Colony collapse disorder (bottom); photo by 21st Centech
Colony collapse disorder
Normal (top), Colony collapse disorder (bottom); photo by 21st Centech

Bees on landing board ventilating a hive; photo by The BeeMD photo collection Bees on landing board ventilating a hive; photo by The BeeMD photo collection
Cooling hive
Bees on landing board ventilating a hive; photo by The BeeMD photo collection
Bee bearding is thought to help cool the bee hive on warm, humid evenings; photo by Mann Lake (Brenda Bray) Bee bearding is thought to help cool the bee hive on warm, humid evenings; photo by Mann Lake (Brenda Bray)
Cooling hive
Bee bearding is thought to help cool the bee hive on warm, humid evenings; photo by Mann Lake (Brenda Bray)
Bees fanning at entrance to cool hive; photo by Dewey M. Caron Bees fanning at entrance to cool hive; photo by Dewey M. Caron
Cooling hive
Bees fanning at entrance to cool hive; photo by Dewey M. Caron
Dark, aged worker comb cells; photo by The BeeMD photo collection Dark, aged worker comb cells; photo by The BeeMD photo collection
Darkening of comb
Dark, aged worker comb cells; photo by The BeeMD photo collection

Aging, darkening brood comb (right), less dark comb from honey (left); photo by Dewey M. Caron Aging, darkening brood comb (right), less dark comb from honey (left); photo by Dewey M. Caron
Darkening of comb
Aging, darkening brood comb (right), less dark comb from honey (left); photo by Dewey M. Caron
Aging, darkening of brood comb; photo by Dewey M. Caron Aging, darkening of brood comb; photo by Dewey M. Caron
Darkening of comb
Aging, darkening of brood comb; photo by Dewey M. Caron
Dark, aged comb; photo by The BeeMD photo collection Dark, aged comb; photo by The BeeMD photo collection
Darkening of comb
Dark, aged comb; photo by The BeeMD photo collection
Queen missing wings; photo by Robert Snyder Queen missing wings; photo by Robert Snyder
Dead or injured queen
Queen missing wings; photo by Robert Snyder

Dead queen outside colony entrance; photo by Evan Fuller Dead queen outside colony entrance; photo by Evan Fuller
Dead or injured queen
Dead queen outside colony entrance; photo by Evan Fuller
Deadout likely caused by PMS; photo by Dewey M. Caron Deadout likely caused by PMS; photo by Dewey M. Caron
Deadouts
Deadout likely caused by PMS; photo by Dewey M. Caron
Deadout: dead bees with their heads in cells (most bees in outer cluster were removed); photo by The BeeMD photo collection Deadout: dead bees with their heads in cells (most bees in outer cluster were removed); photo by The BeeMD photo collection
Deadouts
Deadout: dead bees with their heads in cells (most bees in outer cluster were removed); photo by The BeeMD photo collection
Remains of a dead winter cluster; bee butts fill empty cells in futile attempt to keep warm; photo by The BeeMD photo collection Remains of a dead winter cluster; bee butts fill empty cells in futile attempt to keep warm; photo by The BeeMD photo collection
Deadouts
Remains of a dead winter cluster; bee butts fill empty cells in futile attempt to keep warm; photo by The BeeMD photo collection

Deadout cluster likely due to freezing because of too few bees; photo by Dewey M. Caron Deadout cluster likely due to freezing because of too few bees; photo by Dewey M. Caron
Deadouts
Deadout cluster likely due to freezing because of too few bees; photo by Dewey M. Caron
Deadout in top bar hive; note bees in the bottom of box; photo by Dewey M. Caron Deadout in top bar hive; note bees in the bottom of box; photo by Dewey M. Caron
Deadouts
Deadout in top bar hive; note bees in the bottom of box; photo by Dewey M. Caron
Deadout of a strong colony due to starvation: photo by Kerry Clark Deadout of a strong colony due to starvation: photo by Kerry Clark
Deadouts
Deadout of a strong colony due to starvation: photo by Kerry Clark
Evidence of spring deadout; a dead cluster is with capped brood; photo by Dewey M. Caron Evidence of spring deadout; a dead cluster is with capped brood; photo by Dewey M. Caron
Deadouts
Evidence of spring deadout; a dead cluster is with capped brood; photo by Dewey M. Caron