Endeostigmata

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Taxonomy

Proteonematalycidae Kethley, 1989Kethley, 1989:
Kethley J. 1989. Proteonematalycidae (Acari: Acariformes), a new mite family from fore-dune sand of Lake Michigan. International Journal of Acarology 15 (4): 209-217.

Diagnosis

Elongate endeostigmatans without prodorsal trichobothria, but with eyes and rutella.

Remarks

Proteonematalycus, an even more worm-likeworm-like:
elongate, with reduced legs as in some Nematalycidae (Endeostigmata).
mite, has an unpaired median seta ro and minute prodorsal setae exa, in and exp. It also retains median and lateral eyes, as well as rutellarutellum:
(pl. rutella) In Sarcoptiformes, the hypertrophied setae on the hypostome, often toothed; not to be confused with a corniculus (although possibly a homologue).  In Astigmata, the rutellum may be referred to as a pseudorutellum, although it is the same structure.  Various forms of rutella are recognized, including the atelebasic and pantelebasic.
and 3 pairs of genital papillaegenital papillae:
 1-3 pairs of extrusible finger-like to button-like projections, usually retracted into in the genital vestibule of acariform mites; sometimes formed as sessile disks around the genital opening; thought to be osmoregulatory structures; modified or multiplied and dispersed over the body in many freshwater mites.  Genital papillae are absent in the larva, but may be added ontogenetically: protonymphs have one pair, deutonymphs two pairs, and tritonymphs (and adults) three pairs.  The tritonymphal pair of papillae is often lost.  The serially homologous Claparède's organ is usually present in the larvae (and prelarvae) of mites exhibiting genital papillae in nymphs and adults (Oudeman's Rule).
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References

• Kethley 1989Kethley 1989:
  Kethley J. 1989. Proteonematalycidae (Acari: Acariformes), a new mite family from fore-dune sand of Lake Michigan. International Journal of Acarology 15 (4): 209-217.
• Kethley 1990aKethley 1990a:
  Kethley JB. 1990a. Acarina: Prostigmata (Actinedida). pp. 667-756. In: Dindal DL, ed. Soil biology guide. John Wiley & Sons, Brisbane.