Lissocarpaceae

Disclaimer

Content is from Kirkbride et al. 2006, without modification. 
Updates are forthcoming.

Taxonomy

Lissocarpaceae E. Gilg

Common name: Lissocarpus Family.

Number of genera: 1 genus.

Number of species (Mabberley 1997): 2–3 species.

Disseminule

Fruit (intact or entire), or incomplete fruit with epicarp and mesocarp absent and endocarp exposed.

Description

Fruits: Pistil(s) compound; 1; 1-pistillate; with carpels united. Fruit pericarpium; simple; drupe; without persistent central column; not within accessory organ(s); 1-seeded to more than 1 but less than 10-seeded; 1–2-seeded; from 1–5 cm long; 2.6 cm long; with 4-carpellate; with carpels united; with carpels remaining united at maturity; without sterile carpels; not sulcate; in transection terete; apex not beaked; wall fleshy; indehiscent. Epicarp yellow, or pink (ish); durable; glabrous; without armature; smooth; without wing(s); without apical respiratory hole. Mesocarp present; fleshy; composed of 1 unified layer; without lactiform cavity system; & endocarp sharply differentiated. Endocarp present; not separating from exocarp; thin; not splitting into 1-seeded pyrenes; without operculum; without secretory cavities; without mechanism for seedling escape; without grooves; without longitudinal ridges. Funiculus short; short without seed bearing hooks (retinacula); not persisting in fruit after seed shed.

Seeds: Aril absent. Seed larger than minute; oblong; in transection terete (assumed); not bowl shaped; not nutlike; without winglike beak; without caudate appendage(s); at maturity with food reserves; with endosperm; without canavanine. Sarcotesta absent. Testa present; without markedly different marginal tissue; without fleshy or leathery layer over hard layer; tight (assumed); without crease or line separating cotyledons from hypocotyl-radicle; without notch along margin where cotyledons from hypocotyl-radicle tip approach each other; without glands; without bristles; pubescent; without glandular pubescence; without wings; without collar; without operculum; colored; monochrome; not becoming mucilaginous when wetted; surrounding food reserve. Endosperm copious; hard (horny); smooth; without fatty acid containing cyclopropene; without apical lobes; without chlorophyll; without isodiametric faceted surface; without odor. Embryo differentiated from food reserve; well developed; 1 per seed; partially filling testa (with food reserve); 0.5 times the length of food reserve; at one end of seed not extending into a depression or cup; axile and centric; linear; straight; parallel to seed length; embedded in endosperm; with cotyledons gradually connected to hypocotyl-radicle; without coleorhiza; without simmondsin; without stomata; not green; with 2 or more cotyledons. Cotyledons 2; as wide as hypocotyl-radicle; 1 times wider than hypocotyl-radicle; not concealing hypocotyl-radicle; equal in size; not punctate dotted.

Distribution

General distribution: New World.

Detailed distribution: Tropical South America.

Notes

Spujt did not cover this family.

References

General references: Cronquist, A. 1981. An integrated system of classification of flowering plants, 1,262 p. Columbia University Press, New York, Goldberg, A. 1986 (dicots) & 1989 (monocots). Classification, evolution, and phylogeny of the familes of Dicotyledons. Smithsonian Contr. Bot. 58 for dicots (314 pp.) & 71 for monocots (74 pp.). [Goldberg's illustrations are reproduced from older publications and these should be consulted], Gunn, C.R., J.H. Wiersema, C.A. Ritchie, & J.H. Kirkbride, Jr. 1992 & amendments. Families and genera of Spermatophytes recognized by the Agricultural Research Service. Techn. Bull. U.S.D.A. 1796:1–500, and Mabberley, D.J. 1987. The plant-book, 706 p. Cambridge University Press, Cambridge.

 Fruit:  Lissocarpa benthamii ; Photo by R. Gibbons, USDA APHIS PPQ, Kirkbride et al. (2006)
Fruit: Lissocarpa benthamii; Photo by R. Gibbons, USDA APHIS PPQ, Kirkbride et al. (2006)
 Fruit:  Lissocarpa benthamii ; Photo by R. Gibbons, USDA APHIS PPQ, Kirkbride et al. (2006)
Fruit: Lissocarpa benthamii; Photo by R. Gibbons, USDA APHIS PPQ, Kirkbride et al. (2006)
 Seed:  Lissocarpa benthamii ; Photo by R. Gibbons, USDA APHIS PPQ, Kirkbride et al. (2006)
Seed: Lissocarpa benthamii; Photo by R. Gibbons, USDA APHIS PPQ, Kirkbride et al. (2006)
 Embryo:  Lissocarpa benthamii ; Illustration by K. Parker, Kirkbride et al. (2006)
Embryo: Lissocarpa benthamii; Illustration by K. Parker, Kirkbride et al. (2006)