Emblingiaceae

Disclaimer

Content is from Kirkbride et al. 2006Kirkbride et al. 2006:
Kirkbride JH, Jr, Gunn CR, and Dallwitz MJ. 2006. Family guide for fruits and seeds, vers. 1.0. Accessed September 2020-January 2022. URL: https://nt.ars-grin.gov/seedsfruits/keys/frsdfam/index.cfm .
, without modification. Updates are forthcoming.

Taxonomy

Emblingiaceae (F. Pax) Airy Shaw

Common name: Emblingia Family.

Number of genera: 1 genus.

Number of species (Mabberley 1997): 1 species (Emblingia calecolifora F. Muell.).

Disseminule

Fruit (intact or entire).

Description

Fruits: Pistil(s) 1; 1-pistillate. Fruit without persistent central column; within accessory organ(s); within calyxcalyx:
the outer whorl of the perianth; all the sepals of a flower
(subtending); soft calyxcalyx:
the outer whorl of the perianth; all the sepals of a flower
(soft); without sterilesterile:
lacking male and/or female reproductive parts; also, not producing fruit or seed
carpels; apexapex:
the point farthest from the point of attachment, or the "tip" of an organ
not beaked; wall chartaceouschartaceous:
papery, papyraceous
, or leatheryleathery:
texture—moderately thick, tough, and very pliable
(probably one of these answers); indehiscentindehiscent:
not opening on its own, as in a fruit
(according to literature). Epicarpepicarp:
outer layer of fruit wall or pericarp, if divided into layers; note here used synonymously with exocarp
durable; not glabrousglabrous:
without hairs
(with hairs); hairs not glandularglandular:
surface relief—covered with small, raised secretory glands, regular or irregularly shaped, translucent or opaque, and maybe distinctly colored
; without armature; with wing(s); without apicalapical:
at or pertaining to the end of the seed or fruit distal from its point of attachment (i.e., base)
respiratory hole. Endocarpendocarp:
the inner layer of the pericarp, if divided into layers
present; not separating from exocarpexocarp:
outer layer of fruit wall or pericarp, if divided into layers; note here used synonymously with epicarp
; thin; not splitting into 1-seeded pyrenes; smooth; without wing; without operculumoperculum:
a dehiscent cap (or lid) of a seed or fruit that opens during germination or dehiscence
; without secretory cavities; without mechanism for seedling escape; without grooves; without longitudinallongitudinal:
of or relating to length or the lengthwise dimension
ridges. Funiculusfuniculus:
(alt. funicle) stalk connecting the ovule (later seed) to the ovary (later fruit) placenta
short; short without seed bearing hookswith hooks:
bristles or spines with curved or backwards pointing tips, or with secondary bristles along their length
(retinacula); persisting in fruit after seed shed.

Seeds: Arilaril:
(broad sense) appendicular structure that wholly or partly envelops a seed and is produced from or a modification of the funicle, raphe, or outer integument; usually fleshy or pulpy, sometimes spongy or tufted-capillate, often brightly colored
present (Mueller, 1861, for arillate seed illustration); a true arilaril:
(broad sense) appendicular structure that wholly or partly envelops a seed and is produced from or a modification of the funicle, raphe, or outer integument; usually fleshy or pulpy, sometimes spongy or tufted-capillate, often brightly colored
; fleshy; of funicularfuniculus:
(alt. funicle) stalk connecting the ovule (later seed) to the ovary (later fruit) placenta
origin; marginalmarginal:
at, on, or close to the margin or border
; fimbriate-laciniate (fingerlike projections). Seed circularcircular:
(of embryo) linear embryo is curved into an "O" shape
(more or less); in transection compressedcompressed:
flattened; in grasses, used to denote compression (not necessarily flattened) either laterally or dorsiventrally
; not bowl shaped; not nutlike; without winglike beakbeak:
a usually firm, terminal appendage, sometimes tapered
; without caudatecaudate:
tapering to a long, tail-like appendage
appendage(s); at maturity with food reserves; with endosperm; without canavanine. Sarcotestasarcotesta:
pulpy or fleshy outer layer of the seed coat, simulates aril
absent. Testatesta:
seed coat
absent; adhering to epicarpepicarp:
outer layer of fruit wall or pericarp, if divided into layers; note here used synonymously with exocarp
; without embryo surrounded and capped by viscid tissue. Endosperm scant; smooth; without fatty acid containing cyclopropene; without apicalapical:
at or pertaining to the end of the seed or fruit distal from its point of attachment (i.e., base)
lobes; without chlorophyll; without isodiametric faceted surface; without odor. Embryo differentiated from food reserve; well developed; 1 per seed; nearly filling testatesta:
seed coat
(trace or scanty food reserve); 3 times the length of food reserve (estimated); at one end of seed not extending into a depression or cup; axileaxile:
on or of the axis
and centric; linearlinear:
(shape) long, narrow, and uniform in width; (of embryo) embryo is straight and much longer than wide
; flatly coiledcoiled:
(of embryo) linear embryo is very long and bent to form a coil whereby one end of the embryo is on the outside and the other end near the middle of the seed
(circinatecircinate:
3D shape—terete and rolled downward from the apex in a tight coil
); without coleorhiza; without simmondsin; without stomata; not green; with 2 or more cotyledons. Cotyledons 2; moderately developed; 0.8 times length of embryo; as wide as hypocotyl-radicle; not concealing hypocotyl-radicle; not foliaceous; controtiplicate; smooth; with apicesapex:
the point farthest from the point of attachment, or the "tip" of an organ
entire; with margins separate; basally entire; equal in size; not punctatepunctate:
surface relief—dotted with pits or with translucent, sunken glands or with colored dots, similar to pitted
dotted. Hypocotyl-radicle moderately developed; curvedcurved:
(of embryo) linear embryo is curved into an arch or horseshoe with the ends far apart
; not thickened.

Distribution

General distribution: Old World.

Detailed distribution: Australia.

Notes

Spjut did not cover this family. Airy Shaw: Fruit dry, indehiscentindehiscent:
not opening on its own, as in a fruit
, pendulous within the calyxcalyx:
the outer whorl of the perianth; all the sepals of a flower
from the apexapex:
the point farthest from the point of attachment, or the "tip" of an organ
of the androgynophore, thin pericarppericarp:
fruit wall or fruit coat
adherent to seed. Mueller (1861): Fruit drupaceous, pericarppericarp:
fruit wall or fruit coat
thin.. Calyxcalyx:
the outer whorl of the perianth; all the sepals of a flower
winged.

References

Literature specific to this family: Mueller, F. 1861. Fragmenta phytographie Australiae 2:1–4 & plate 11.

General references: Airy Shaw, H.K. 1973. A dictionary of the flowering plants and ferns, 1,131 pp. University Press, Cambridge, Engler, A. & K. Prantl. 1924 and onward. Die Natürlichen Pflanzenfamilimien. W. Engelman, Leipzig, Gunn, C.R., J.H. Wiersema, C.A. Ritchie, & J.H. Kirkbride, Jr. 1992 & amendments. Families and genera of Spermatophytes recognized by the Agricultural Research Service. Techn. Bull. U.S.D.A. 1796:1–500, and Mabberley, D.J. 1987. The plant-book, 706 p. Cambridge University Press, Cambridge.

 Embryo:    Emblingia calceoliflora ; Illustration by K. Parker, Kirkbride et al. (2006)
Embryo: Emblingia calceoliflora; Illustration by K. Parker, Kirkbride et al. (2006)