Authors and acknowledgments

Authors

Nathan P. Lord, Ph.D., was an assistant professor at Georgia College & State University. Nathan is broadly interested in Coleoptera systematics and has published works on several families, including Bothrideridae, Cerambycidae, Endomychidae, Latridiidae, and Zopheridae. Visit Nathan's Coleoptera Systematics website.

Eugenio H. Nearns, Ph.D., was a Coleoptera specialist with APHIS/PPQ's National Identification Services (NIS) (2017-2025). Gino's research is focused on the taxonomy, systematics, and evolution of longhorned beetles. Gino has described >73 new species and published >70 papers, including a revision of the cerambycine tribes Curiini and Plectromerini (Nearns and Branham 2008Nearns and Branham 2008:
Nearns EH, Branham MA. 2008. Revision and phylogeny of the tribes Curiini Le-Conte and Plectromerini Nearns & Branham, new tribe (Coleoptera: Cerambyci-dae: Cerambycinae). Memoirs of the American Entomological Society. 47:1–117.) and a Lucid key to the 80 genera of lamiine tribe Onciderini (Nearns et al. 2011Nearns et al. 2011:
Nearns EH, Lord NP, Miller KB. 2011. Oncid ID: Tool for diagnosing adult twig girdlers (Cerambycidae: Lamiinae: Onciderini). The University of New Mexico and Center for Plant Health Science and Technology, USDA, APHIS, PPQ.). Visit Gino's Cerambycid Research website.

Kelly B. Miller, Ph.D., is an associate professor at the Department of Biology and curator of Arthropods at the Museum of Southwestern Biology, University of New Mexico, Albuquerque, NM, USA. Kelly has worked extensively at the alpha-taxonomic level revising genera and species groups, and has also conducted higher-taxonomic phylogenetic work using cladistic analyses of morphological and molecular data to improve classifications and examine evolutionary scenarios. Visit the Kelly Miller Lab of Insect Systematics website.

Website migration

Robin Boudwin, Center for Integrated Pest Management, North Carolina State University, Raleigh, North Carolina
Melissa Islam, Department of Biology, Colorado State University, Fort Collins, Colorado
Ashley Gottlieb, Department of Biology, Colorado State University, Fort Collins, Colorado
Kojun Kanda, USDA-APHIS-PPQ-S&T, Pest Identification Technology Lab (PITL), Identification Technology Program (ITP), Washington, DC

In 2026, tool content was migrated to this website created using Fact Sheet Manager 3, a content-management system, developed and maintained by Center for Integrated Pest Management, North Carolina State University.

For questions about content, access, or functionality, please contact ITP (itp@usda.gov).

Acknowledgments

The authors would like to thank Terrence Walters and Amanda Redford, USDA/APHIS/PPQ/CPHST/ITP and the University of New Mexico, Department of Biology, for funding the development of Ironclad ID.

We thank Richard A.B. Leschen, Paul E. Skelley, and James E. Zablotny for critical suggestions to an earlier version of this tool. Helpful comments and suggestions were also provided by Grey T. Gustafson and Dr. Kelly Miller's Biology 485/585 Entomology class (University of New Mexico, Albuquerque, NM).

We are especially grateful to the following individuals for facilitating our collections-based research: Max Barclay, Sharon Shute, and Roger Booth (The Natural History Museum), Steven W. Lingafelter (Systematic Entomology Lab / US National Museum), Michael C. Thomas and Paul E. Skelley (Florida State Collection of Arthropods), Thierry Deuve and Azadeh Taghavian (Muséum National d'Histoire Naturelle), Adam Slipinski (CSIRO Entomology, Canberra, Australia), and Richard Leschen (New Zealand Collection of Arthropods).

The following individuals generously loaned specimens and provided support: Thierry Deuve and Azadeh Taghavian (Muséum National d'Histoire Naturelle, Paris, France), Steve Lingafelter, Norm Woodley, Warren Steiner, Gary Hevel and David Furth (US National Museum/Smithsonian Institution), Paul Skelley and Mike Thomas (FSCA, Gainesville, FL), Max Barclay and Roger Booth (BMNH, London, UK), Adam Slipinski (CSIRO Entomology, Canberra, Australia), Mike Ulyshen (UGCA, Athens, GA), Dave Lightfoot and Sandy Brantley (MSB, Albuquerque, NM), Andy Cline (CDFA, Sacramento, CA), Rich Leschen (Landcare Research, Auckland, New Zealand), Mike Ivie (Montana State University, Bozeman, MT), Joe McHugh (University of Georgia, Athens, GA), Ken Karns (Columbus, Ohio), Richard Buss (Albuquerque, NM), and Heather L. Paulsen (University of New Mexico).

Roy Larimer (Visionary Digital) provided support for the BK Plus and Passport imaging systems.

Matt Taylor and Damian Barnier (Lucidcentral.org) provided software support.