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CAPS Secondary Target - Adult

Archips fuscocupreanus Walsingham (Tortricidae: Tortricinae: Archipini)

Common names: apple tortrix

Synonyms: ishidaii (Loxotaenia), punicae (Cacoecia), rosaceana (Archips)

Fig. 1: Male

Fig. 1: Male

Fig. 2: Female

Fig. 2: Female

Fig. 3: Female

Fig. 3: Female

Fig. 4: Male genitalia

Fig. 4: Male genitalia

Fig. 5: Female genitalia

Fig. 5: Female genitalia

Adult Recognition

FWL: 7.0-9.5 mm (male); 8.0-12.5 mm (female)

Forewing color is dull brown with dark-brown to reddish-brown markings. Pattern expression can vary between individuals, but the median fascia is complete from costa to dorsum. Hindwings are brownish gray. Males have a forewing costal fold.

Some individuals resemble Archips xylosteana, but the two species can be separated by the broader costal fold and continuous median fasica (from costa to dorsum) in A. fuscocupreanus.

Other Archips species, including Archips crataegana, Archips grisea, and forms of Archips podana and Archips rosana, can appear similar to A. fuscocupreanus. A genitalic dissection can be used to confirm identity.

Larval Morphology

Late instar larvae are approximately 22 mm in length. The abdomen is pale green to grayish green with dark dorsal and lateral lines and conspicuous pale pinacula. The head is orange or brownish orange. The prothoracic shield is brownish orange with black posterolateral margins. The thoracic legs and thoracic pinacula are black.


Archips fuscocupreanus completes one generation per year, with adults present in mid-June through mid-July in the northeastern U.S.

Females lay black egg masses on the trunks and limbs of trees. The eggs overwinter, and first instar larvae hatch the following spring. Early instars feed on young leaves. Later instars construct a shelter by webbing together leaves and consume leaves, flowers, and occasionally fruit. Pupation occurs in webbed leaves.

Host plants

Larvae of Archips fuscocupreanus are highly polyphagous and have been recorded feeding on plants in more than 23 families. Many larval hosts are in the Rosaceae, and A. fuscocupreanus is an important pest of apple in East Asia. A partial host list is presented below; Maier (2003) provides a list of other minor hosts found in the northeastern U.S.

Family Genus/species Common name
Aceraceae Acer L. maple
Aquifoliaceae Ilex verticillata (L.) Gray common winterberry
Asteraceae Erigeron annuus (L.) Pers. eastern daisy
Betulaceae Alnus Mill. alder
Betulaceae Betula platyphylla Sukaczev Asian white birch
Betulaceae Corylus L. hazelnut
Celastraceae Celastrus orbiculatus Thunb. Oriental bittersweet
Cornaceae Cornus L. dogwood
Ebenaceae Diospyros kaki L. f. Japanese persimmon
Elaeagnaceae Elaeagnus umbellata Thunb. autumn olive
Ericaceae Rhododendron L. rhododendron
Euphorbiaceae Ricinus communis L. castorbean
Fabaceae Glycine max (L.) Merr. soybean
Fabaceae Phaseolus L. bean
Fagaceae Castanea crenata Siebold & Zucc. Japanese chestnut
Fagaceae Castanea Mill. chestnut
Fagaceae Quercus acutissima Carruth. sawtooth oak
Fagaceae Quercus glauca Thunb. ring-cup oak
Fagaceae Quercus mongolica Fisch. ex Ledeb. mongolian oak
Fagaceae Quercus serrata Thunb. bao li
Fagaceae Quercus variabilis Blume Chinese cork oak
Hydrangeaceae Deutzia Thunb. pride-of-Rochester
Juglandaceae Juglans L. walnut
Moraceae Morus alba L. white mulberry
Moraceae Morus bombycis Koidz.
Pinaceae Larix Mill. larch
Pinaceae Picea A. Dietr. spruce
Rhamnaceae Rhamnus L. buckthorn
Rosaceae Amelanchier canadensis (L.) Medik. Canadian serviceberry
Rosaceae Cydonia oblonga Mill. quince
Rosaceae Fragaria L. strawberry
Rosaceae Malus baccata (L.) Borkh. Siberian crab apple
Rosaceae Malus pumila Mill. paradise apple
Rosaceae Malus Mill. apple
Rosaceae Prunus armeniaca L. apricot
Rosaceae Prunus avium (L.) L. sweet cherry
Rosaceae Prunus laurocerasus L. cherry laurel
Rosaceae Prunus mume Siebold & Zucc. Japanese apricot
Rosaceae Prunus persica (L.) Batsch peach
Rosaceae Prunus salicina Lindl. Japanese plum
Rosaceae Prunus sargentii Rehder
Rosaceae Prunus serotina Ehrh. black cherry
Rosaceae Prunus X yedoensis Matsum. (pro sp.) [subhirtella X speciosa]
Rosaceae Pyrus pyrifolia (Burm. f.) Nakai Chinese pear
Rosaceae Pyrus ussuriensis Maxim.
Rosaceae Pyrus L. pear
Rosaceae Rosa multiflora Thunb. multiflora rose
Rosaceae Rosa rugosa Thunb. rugosa rose
Rosaceae Rubus L. blackberry
Rosaceae Sorbus L. mountain ash
Salicaceae Salix L. willow
Ulmaceae Ulmus L. elm


A native of East Asia, Archips fuscocupreanus has been introduced to the northeastern United States (Connecticut, Massachusetts, New Jersey, New York, and Rhode Island) and Pacific Northwest (Washington).


Maier, C. T. 2003. Distribution, hosts, abundance, and seasonal flight activity of the exotic leafroller, Archips fuscocupreanus Walsingham (Lepidoptera: Tortricidae), in the northeastern United States. Annals of the Entomological Society of America. 96: 660-666.

Maier, C. T. and V. C. Mastro. 1998. Discovery, abundance, and distribution of the exotic apple tortrix, Archips fuscocupreanus Walsingham (Lepidoptera: Tortricidae), in the northeastern United States. Proceedings of the Entomological Society of Washington. 100: 545-552.

Yasuda, T. 1975. The Tortricinae and Sparganothinae of Japan (Lepidoptera: Tortricidae). Part II. Bulletin of the University of Osaka Prefecture, Series B. 27: 79-251.

Tortricids of Agricultural Importance by Todd M. Gilligan and Marc E. Epstein
Interactive Keys developed in Lucid 3.5. Last updated August 2014.