Asparagaceae

Taxonomy

Family name: Asparagaceae Juss.

Synonym(s): Agavaceae Dumort., nom. cons.; Anemarrhenaceae Conran et al.; Anthericaceae J. Agardh; Aphyllanthaceae Burnett; Aspidistraceae Hassk.; Behniaceae Conran et al.; Convallariaceae Horan.; Dracaenaceae Salisb., nom. cons.; Eriospermaceae Lem.; Eucomidaceae Salisb.; Eustrephaceae Chupov; Funkiaceae Horan., nom. illeg.; Herreriaceae Kunth; Hesperocallidaceae Traub; Hostaceae B. Mathew; Hyacinthaceae Batsch ex Borkh.; Lachenaliaceae Salisb.; Laxmanniaceae Bubani; Lomandraceae Lotsy; Nolinaceae Nakai; Ophiopogonaceae Meisn.; Ornithogalaceae Salisb.; Platymetraceae Salisb., nom. illeg.; Polygonataceae Salisb.; Ruscaceae M. Roem., nom. cons.; Sansevieriaceae Nakai; Scillaceae Vest; Themidaceae Salisb.; Xerotaceae Hassk., nom. illeg.; Yuccaceae J. Agardh;

Common name(s): asparagus family

*Number of genera/species: 114/2900

List of genera records in GRIN-Global

Disseminule

fruit or seed

Description

Fruit indehiscent, berryberry:
an indehiscent, fleshy fruit with one or a few to many seeds. The flesh may be homogenous throughout. Or, if the outer part is hard, firm, or leathery, referred to as an hesperidium. Septa are present in some, and the seeds may be arillate or with a fleshy testa.
or dehiscent, loculicidalloculicidal:
type of capsular dehiscence, opening longitudinally through the locules (compare septicidal)
or septicidalsepticidal:
type of capsular dehiscence, opening longitudinally by separating between the septa of adjacent carpels
capsulecapsule:
a dry, dehiscent fruit formed by two or more carpels
, or rarely a schizocarpschizocarp:
fruit formed from a single ovary, with fused carpels, with or without accessory tissue; splitting between locules to form distinct, indehiscent, usually one seeded segments; usually dry, rarely fleshy (compare mericarp)
(Ledebouria), 2–230 mm long, globoseglobose:
3D shape—more or less spherical
to angularangular:
2D shape—having sides that meet at acute or obtuse angles  
, sometimes sulcatesulcate:
surface relief—having one or more elongate, relatively narrow and shallow depressions or grooves
, tereteterete:
approximately circular in cross section; width and thickness approximately equal
or angledangled:
2D shape—having sides that meet at acute or obtuse angles  
in transectiontransection:
a cross section; representing a plane made by cutting across an organ at a right angle to its length
, sometimes with a persistent stylestyle:
in a flower, the narrow and elongated part of the pistil between the stigma and the ovary; sometimes persisting in fruit
or perianthperianth:
collective term for calyx and corolla of a flower
, one seeded or up to 10 seeded. Pericarppericarp:
fruit wall or fruit coat
black or brown (capsules) or red, orange, blue, purple, or black (berries), shinyshiny:
uniformly reflecting a high proportion of incident light at all angles
or dulldull:
reflecting only a low proportion of incident light, with no apparent sheen
, fleshyfleshy:
texture—fairly firm and dense, juicy or at least moist, and easily cut
to chartaceouschartaceous:
=papery, papyraceous
, glabrousglabrous:
without hairs
or pubescentpubescent:
surface relief—bearing hairs
, smooth, ribbed, wartywarty:
surface relief—distinct, rounded projections that are large relative to the fruit size; tuberculate, verrucose
, or spinyspiny:
having slender, stiff, sharp projections oriented in the general plane of the structure
. Fruits in Nolina and some species of Chlorophytum are three-winged. Agave species have beakedbeaked:
a usually firm, terminal appendage, sometimes tapered
fruits. Eriospermum’s loculicidalloculicidal:
type of capsular dehiscence, opening longitudinally through the locules (compare septicidal)
capsulescapsules:
a dry, dehiscent fruit formed by two or more carpels
(globose or lobed) have unique long, unicellular hairs up to 8 mm long.

Seed globoseglobose:
3D shape—more or less spherical
to angledangled:
2D shape—having sides that meet at acute or obtuse angles  
, including discoidaldiscoidal:
3D shape—resembling a disc
to irregular, tereteterete:
approximately circular in cross section; width and thickness approximately equal
, angledangled:
2D shape—having sides that meet at acute or obtuse angles  
or flattened in transectiontransection:
a cross section; representing a plane made by cutting across an organ at a right angle to its length
, 0.7–20 mm long. Liriope and Ophiopogon seeds have bright, blue sarcotestasarcotesta:
pulpy or fleshy outer layer of the seed coat, simulates aril
. Seed coat black or brown, rarely orange, red, or white, hard and thinthin:
having or being of relatively little depth
, shinyshiny:
uniformly reflecting a high proportion of incident light at all angles
or dulldull:
reflecting only a low proportion of incident light, with no apparent sheen
, smooth, wrinkled, pittedpitted:
surface relief—surface with small depressions in which the areas between the hollows do not take on the appearance of a true reticular net
, papillatepapillate:
surface relief—bearing minute, distinct, broad-based projections, tapering to a rounded apex
, or reticulatereticulate:
surface relief—netted, raised walls or concave grooves forming a net-like surface pattern with flat, concave, or convex interspaces
, and sometimes pubescentpubescent:
surface relief—bearing hairs
. Seeds in capsular fruit have phytomelanphytomelan:
carbonaceous, opaque material that usually covers the seed coat to give it a black appearance, common in certain monocot families
except in Eriospermum.

An elaiosomeelaiosome:
a lipid and protein-rich fleshy structure attached to some seeds and fruits, it attracts ants which then disperse the disseminule (e.g., caruncle in the Euphorbiaceae, the aril (outgrowth of the funiculus) in the Fabaceae)
is present in some Caesia, Hensmania, and Stawellia species; a white carunclecaruncle:
a localized outgrowth of the seed coat near the hilum of the seed; it functions as an elaiosome
is present in Johnsonia (seed black and reniformreniform:
2D or 3D shape—kidney-shaped
). Seeds are helically winged in Herreria and Herreriopsis.

Embryo well developed, linearlinear:
(shape) long, narrow, and uniform in width; (of embryo) embryo is straight and much longer than wide
, cylindric or broad, straight or curvedcurved:
(of embryo) linear embryo is curved into an arch or horseshoe with the ends far apart
(Anthericum, Arthropodium, Chlorogalum, Dichopogon, and Thysanotus), axile and centric. Endosperm copious, cartilaginouscartilaginous:
texture—firm, dense, tough, somewhat pliable, and resilient, like cartilage
or hard, smooth or corrugated.

Identification features

Fruit
Type berry, loculicidalloculicidal:
type of capsular dehiscence, opening longitudinally through the locules (compare septicidal)
or septicidalsepticidal:
type of capsular dehiscence, opening longitudinally by separating between the septa of adjacent carpels
capsulecapsule:
a dry, dehiscent fruit formed by two or more carpels
, rarely schizocarp
Size range 2–230 mm long
Shape(s) globose, ovoidovoid:
3D shape—ovate
, oblongoblong:
2D shape—much longer than broad with nearly parallel sides, corners are rounded
, cylindricalcylindrical:
3D shape—a cylinder, with parallel sides and a circular cross-section; tubular or rod-shaped
, conicalconical:
3D shape—cone-shaped, with the point of attachment at the broad end
, trigonoustrigonous:
3D shape—having three faces that meet at distinct angles; triangular in outline
, triangulartriangular:
2D shape—three relatively straight sides with distinct corners; more angular than teardrop-shaped
, angled
Texture epicarp epicarp:
outer layer of fruit wall or pericarp, if divided into layers; note here used synonymously with exocarp
- fleshy, spongyspongy:
soft, light, discontinuous but cohesive, and somewhat resilient
, leatheryleathery:
texture—moderately thick, tough, and very pliable
, chartaceouschartaceous:
=papery, papyraceous
; mesocarpmesocarp:
the middle layer of the pericarp, if divided into layers
- drydry:
texture—free or relatively free from water or liquid
or fleshy
Surface relief smooth or ribbedribbed:
surface relief—wide, prominent, linear ridges that are generally rounded and longitudinally situated on the surface
, spinyspiny:
having slender, stiff, sharp projections oriented in the general plane of the structure
, warty
Color(s) red, orange, blue, purple, black, brown
Unique features Black or brown capsulescapsules:
a dry, dehiscent fruit formed by two or more carpels
often with flattened, disc-shaped or angularangular:
2D shape—having sides that meet at acute or obtuse angles  
seeds. Or, brightly colored to black berriesberries:
an indehiscent, fleshy fruit with one or a few to many seeds. The flesh may be homogenous throughout. Or, if the outer part is hard, firm, or leathery, referred to as an hesperidium. Septa are present in some, and the seeds may be arillate or with a fleshy testa.
with tereteterete:
approximately circular in cross section; width and thickness approximately equal
to angledangled:
2D shape—having sides that meet at acute or obtuse angles  
seeds. In Eriopspermum, capsules with long hairs, up to 8 mm long.
Seed
Size range 0.7–20 mm long
Shape(s) hemispherical, globoseglobose:
3D shape—more or less spherical
, oblongoblong:
2D shape—much longer than broad with nearly parallel sides, corners are rounded
, ovoidovoid:
3D shape—ovate
, ellipsoidellipsoid:
3D shape—elliptic
, cuneiformcuneiform:
=wedge-shaped
, discoidaldiscoidal:
3D shape—resembling a disc
, irregular, reniformreniform:
2D or 3D shape—kidney-shaped
, conicalconical:
3D shape—cone-shaped, with the point of attachment at the broad end
, teardrop-shapedteardrop-shaped:
2D shape—widest point is toward one end of the fruit, the other end tapers sharply to a pointed end
, fusiformfusiform:
spindle-shaped; broadest at the middle and tapering at both ends
, triangulartriangular:
2D shape—three relatively straight sides with distinct corners; more angular than teardrop-shaped
, polygonal
Surface relief wrinkled, papillatepapillate:
surface relief—bearing minute, distinct, broad-based projections, tapering to a rounded apex
, pittedpitted:
surface relief—surface with small depressions in which the areas between the hollows do not take on the appearance of a true reticular net
, reticulate
Color(s) black, brown, orange, red, blue, white
Unique features Seeds with phytomelanphytomelan:
carbonaceous, opaque material that usually covers the seed coat to give it a black appearance, common in certain monocot families
in most genera with capsular fruits. Seeds with bright blue sarcotestasarcotesta:
pulpy or fleshy outer layer of the seed coat, simulates aril
in Liriope and Ophiopogon. Seeds in some genera with arilaril:
(broad sense) appendicular structure that wholly or partly envelops a seed and is produced from or a modification of the funicle, raphe, or outer integument; usually fleshy or pulpy, sometimes spongy or tufted-capillate, often brightly colored
or aril-like structures. Seeds helically winged in two genera. 
Other
Embryo well developed, axileaxile:
on or of the axis
and centric in position, linearlinear:
(shape) long, narrow, and uniform in width; (of embryo) embryo is straight and much longer than wide
, cylindriccylindric:
3D shape—a cylinder, with parallel sides and a circular cross-section; tubular or rod-shaped
, or broad in shape, and straight or curved
Nutritive tissuenutritive tissue:
tissue within the seeds that nourishes the developing embryo; such as endosperm, perisperm, or chalazosperm in angiosperms; megagametophyte in gymnosperms
endosperm endosperm:
nutritive starch- and oil-containing tissue present in many seeds
copious, cartilaginouscartilaginous:
texture—firm, dense, tough, somewhat pliable, and resilient, like cartilage
or hard, smooth or corrugated

Distribution

World-wide, except Arctic.

Distribution map courtesy of Angiosperm Phylogeny Website.

References

Baskin and Baskin 2021Baskin and Baskin 2021:
Baskin C and Baskin J. 2021. Relationship of the lateral embryo (in grasses) to other monocot embryos: A status up-grade. Seed Science Research 31 (3): 199-210. doi:10.1017/S0960258521000209
; Dahlgren et al. 1985Dahlgren et al. 1985:
Dahlgren RMT, Clifford HT, and Yeo PF. 1985. The families of the monocotyledons: structure, evolution, and taxonomy. Springer-Verlag, Berlin. 520 pp.
; Damen et al. 2018Damen et al. 2018:
Damen THJ, van der Burg WJ, Wiland-Szymańska J, and Sosef MSM. 2018. Taxonomic novelties in African Dracaena (Dracaenaceae). Blumea: Journal of Plant Taxonomy and Plant Geography 63 (1): 31-53. http://dx.doi.org/10.3767/blumea.2018.63.01.05
; Davidse et al. 2009–2018Davidse et al. 2009–2018:
Davidse GM, Sousa Sánchez M, Knapp S. and Chiang Cabrera F, eds. 2009–2018. Flora Mesoamericana. Missouri Botanical Garden, St. Louis, MO. Accessed: January–April 2024. URL: http://legacy.tropicos.org/Project/FM
; Demissew 2006Demissew 2006:
Demissew S. 2006. Asparagaceae. In: Beentje HJ and Ghazanfar SA, eds. Flora of Tropical East Africa. Vol 16a. Kew, Richmond UK. 23 pp.
; Flora of Australia 2021+Flora of Australia 2021+:
Flora of Australia. Australian Biological Resources Study, Canberra. Accessed January 2021–March 2024. URL: http://www.ausflora.org.au
; Flora of North America Editorial Committee 1993+Flora of North America Editorial Committee 1993+:
Flora of North America Editorial Committee, eds. 1993+. Flora of North America North of Mexico [Online]. 22+ vols. Flora of North America Association, New York and Oxford. Accessed January-March 2024. URL: http://beta.floranorthamerica.org.
; Kirkbride et al. 2006Kirkbride et al. 2006:
Kirkbride JH, Jr, Gunn CR, and Dallwitz MJ. 2006. Family guide for fruits and seeds, vers. 1.0. Accessed September 2020-January 2022. URL: https://nt.ars-grin.gov/seedsfruits/keys/frsdfam/index.cfm .
; Kubitzki et al. 1990+Kubitzki et al. 1990+:
Kubitzki K et al., eds. 1990+. The families and genera of vascular plants. 7+ vols. Berlin etc.
; Snijman 2013Snijman 2013:
Snijman DA. 2013. Plants of the Greater Cape Floristic Region, Vol 2: The Extra Cape flora. Strelitzia 30. South African National Biodiversity Institute, Pretoria.
; Stevens 2001+Stevens 2001+:
Stevens PF. 2001+. Angiosperm Phylogeny Website Version 14, July 2017 [and more or less continuously updated since]. Accessed September 2020-2022. URL: http://www.mobot.org/MOBOT/research/APweb/
; Stevenson and Loconte 1995Stevenson and Loconte 1995:
Stevenson DW and Loconte H. 1995. A cladistic analysis of monocot families. In: Rudall PJ, Cribb PJ, Cutler DF, and Humphries CJ, eds. Monocotyledons: Systematics and Evolution. Royal Botanic Gardens, Kew.
; Tutin et al. 1964–1980Tutin et al. 1964–1980:
Tutin TG, Burges NA, Chater AO, Edmondson JR, Heywood VH, Moore DM, Valentine DH, Walters SM, and Webb DA (eds.) 19641980. Flora Europaea. 5 vols. Cambridge University Press, Cambridge UK. 2,524 pp.
; Vislobokov et al. 2013Vislobokov et al. 2013:
Vislobokov NA, Kuznetsov AN, and Sokoloff DD. 2013. A new species of Aspidistra (Ruscaceae s.l., Asparagales) from southern Vietnam, field observations on its flowering and possible pollination by flies (Phoridae). Plant Systematics and Evolution 299 (2): 347-355. http://dx.doi.org/10.1007/s00606-012-0725-2
; Watson and Dallwitz 1992+Watson and Dallwitz 1992+:
Watson L and Dallwitz MJ. 1992+. The families of flowering plants: descriptions, illustrations, identification, and information retrieval. Version: 6th Accessed September 2020-September 2022. URL: delta-intkey.com
; Zhengyi et al. 2004+Zhengyi et al. 2004+:
Zhengyi W, Raven PH, and Deyuan H. 2004+. Flora of China [online]. 25 vols. Science Press, Beijing China & Missouri Botanical Garden, St. Louis USA. Accessed January–March 2024. http://flora.huh.harvard.edu/china/

*The number of genera and species is based on Christenhusz and Byng 2016Christenhusz and Byng 2016:
Christenhusz MJM and Byng JW. 2016. The number of known plant species in the world and its annual increase. Phytotaxa 261 (3): 201-217. https://doi.org/10.11646/phytotaxa.261.3.1
, which may differ from the number of genera in GRIN-Global.

  Fruit:   Agave cerulata ; Photo by R. Gibbons, USDA APHIS PPQ, Kirkbride et al. (2006)

Fruit: Agave cerulata; Photo by R. Gibbons, USDA APHIS PPQ, Kirkbride et al. (2006)

  Fruit, seed :  Agave  sp.; Photo by N. Diaz, USDA APHIS PPQ, imageID.idtools.org

Fruit, seed: Agave sp.; Photo by N. Diaz, USDA APHIS PPQ, imageID.idtools.org

  Seeds:   Agave cerulata ; Photo by R. Gibbons, USDA APHIS PPQ, Kirkbride et al. (2006)

Seeds: Agave cerulata; Photo by R. Gibbons, USDA APHIS PPQ, Kirkbride et al. (2006)

  Fruits :  Asparagus aethiopicus ; Photo by A. Yee, flickr.com

Fruits: Asparagus aethiopicus; Photo by A. Yee, flickr.com

  Fruit, seed:   Asparagus officinalis ; Photo by D. Walters and C. Southwick, Table Grape Weed Disseminule ID

Fruit, seed: Asparagus officinalis; Photo by D. Walters and C. Southwick, Table Grape Weed Disseminule ID

  Seeds :  Asparagus officinalis ; Photo by D. Walters and C. Southwick, Table Grape Weed Disseminule ID

Seeds: Asparagus officinalis; Photo by D. Walters and C. Southwick, Table Grape Weed Disseminule ID

  Seeds:   Asparagus officinalis ; Photo by D. Walters and C. Southwick, Table Grape Weed Disseminule ID

Seeds: Asparagus officinalis; Photo by D. Walters and C. Southwick, Table Grape Weed Disseminule ID

  Fruit:   Ornithogalum umbellatum ; Photo by L.J. Mehrhoff, University of Connecticut, bugwood.org

Fruit: Ornithogalum umbellatum; Photo by L.J. Mehrhoff, University of Connecticut, bugwood.org

  Seed:   Ornithogalum  sp.; Photo by A. Margina, USDA APHIS PPQ, imageID.idtools.org
Seed: Ornithogalum sp.; Photo by A. Margina, USDA APHIS PPQ, imageID.idtools.org
  Embryo:   Ornithogalum pyrenaicum ; Illustration by K. Parker, Kirkbride et al. (2006)

Embryo: Ornithogalum pyrenaicum; Illustration by K. Parker, Kirkbride et al. (2006)

  Infructescence :  Lomandra multiflora ; Photo by South Australian Seed Conservation Centre, used with permission

Infructescence: Lomandra multiflora; Photo by South Australian Seed Conservation Centre, used with permission

  Infructescence :  Lomandra longifolia ; Photo by South Australian Seed Conservation Centre, used with permission

Infructescence: Lomandra longifolia; Photo by South Australian Seed Conservation Centre, used with permission

  Fruit, seed :  Lomandra leucocephala ; Photo by South Australian Seed Conservation Centre, used with permission

Fruit, seed: Lomandra leucocephala; Photo by South Australian Seed Conservation Centre, used with permission

  Seeds :  Lomandra fibrata ; Photo by South Australian Seed Conservation Centre, used with permission

Seeds: Lomandra fibrata; Photo by South Australian Seed Conservation Centre, used with permission

  Seeds :  Lomandra densiflora ; Photo by South Australian Seed Conservation Centre, used with permission

Seeds: Lomandra densiflora; Photo by South Australian Seed Conservation Centre, used with permission

  Fruits:   Convallaria majalis ; Photo by A. Oommen, bugwood.org

Fruits: Convallaria majalis; Photo by A. Oommen, bugwood.org

  Seeds :  Convallaria majalis ; Photo by D. Walters and C. Southwick, USDA APHIS PPQ ITP, imageID.idtools.org

Seeds: Convallaria majalis; Photo by D. Walters and C. Southwick, USDA APHIS PPQ ITP, imageID.idtools.org

  Seeds :  Convallaria majalis ; Photo by D. Walters and C. Southwick, USDA APHIS PPQ ITP, imageID.idtools.org

Seeds: Convallaria majalis; Photo by D. Walters and C. Southwick, USDA APHIS PPQ ITP, imageID.idtools.org

  Fruit:   Yucca elata ; Photo by R. Gibbons, USDA APHIS PPQ, Kirkbride et al. (2006)

Fruit: Yucca elata; Photo by R. Gibbons, USDA APHIS PPQ, Kirkbride et al. (2006)

  Seeds:   Yucca elata ; Photo by R. Gibbons, USDA APHIS PPQ, Kirkbride et al. (2006)

Seeds: Yucca elata; Photo by R. Gibbons, USDA APHIS PPQ, Kirkbride et al. (2006)

  Embryo:   Yucca elata ; Illustration by K. Parker, Kirkbride et al. (2006)

Embryo: Yucca elata; Illustration by K. Parker, Kirkbride et al. (2006)

  Fruit:   Muscari muscarimi ; Photo by R. Gibbons, USDA APHIS PPQ, Kirkbride et al. (2006)

Fruit: Muscari muscarimi; Photo by R. Gibbons, USDA APHIS PPQ, Kirkbride et al. (2006)

  Seeds:   Muscari muscarimi ; Photo by R. Gibbons, USDA APHIS PPQ, Kirkbride et al. (2006)

Seeds: Muscari muscarimi; Photo by R. Gibbons, USDA APHIS PPQ, Kirkbride et al. (2006)

  Embryo:   Muscari comosum ; Illustration by K. Parker, Kirkbride et al. (2006)

Embryo: Muscari comosum; Illustration by K. Parker, Kirkbride et al. (2006)

  Fruit, seed:   Cordyline cannifolia ; Photo by R. Gibbons, USDA APHIS PPQ, Kirkbride et al. (2006)

Fruit, seed: Cordyline cannifolia; Photo by R. Gibbons, USDA APHIS PPQ, Kirkbride et al. (2006)

  Fruit:   Paradisea liliastrum ; Photo by R. Gibbons, USDA APHIS PPQ, Kirkbride et al. (2006)

Fruit: Paradisea liliastrum; Photo by R. Gibbons, USDA APHIS PPQ, Kirkbride et al. (2006)

  Seeds:   Paradisea liliastrum ; Photo by R. Gibbons, USDA APHIS PPQ, Kirkbride et al. (2006)

Seeds: Paradisea liliastrum; Photo by R. Gibbons, USDA APHIS PPQ, Kirkbride et al. (2006)

  Fruits:   Acanthocarpus preissii ; Photo by R. Gibbons, USDA APHIS PPQ, Kirkbride et al. (2006)

Fruits: Acanthocarpus preissii; Photo by R. Gibbons, USDA APHIS PPQ, Kirkbride et al. (2006)

  Seeds:   Acanthocarpus preissii ; Photo by R. Gibbons, USDA APHIS PPQ, Kirkbride et al. (2006)

Seeds: Acanthocarpus preissii; Photo by R. Gibbons, USDA APHIS PPQ, Kirkbride et al. (2006)

  Fruit:   Beaucarnea guatemalensis ; Photo by R. Gibbons, USDA APHIS PPQ, Kirkbride et al. (2006)

Fruit: Beaucarnea guatemalensis; Photo by R. Gibbons, USDA APHIS PPQ, Kirkbride et al. (2006)

  Fruits :  Danae racemosa ; Photo by K.E. Clancy, USDA APHIS PPQ, imageID.idtools.org

Fruits: Danae racemosa; Photo by K.E. Clancy, USDA APHIS PPQ, imageID.idtools.org

  Fruits :  Danae racemosa ; Photo by K.E. Clancy, USDA APHIS PPQ, imageID.idtools.org

Fruits: Danae racemosa; Photo by K.E. Clancy, USDA APHIS PPQ, imageID.idtools.org

  Fruit, seed :  Danae racemosa ; Photo by K.E. Clancy, USDA APHIS PPQ, imageID.idtools.org

Fruit, seed: Danae racemosa; Photo by K.E. Clancy, USDA APHIS PPQ, imageID.idtools.org

  Seeds :  Arthropodium fimbriatum ; Photo by South Australian Seed Conservation Centre, used with permission

Seeds: Arthropodium fimbriatum; Photo by South Australian Seed Conservation Centre, used with permission

  Seeds :  Arthropodium strictum ; Photo by South Australian Seed Conservation Centre, used with permission

Seeds: Arthropodium strictum; Photo by South Australian Seed Conservation Centre, used with permission