Immanus
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Immanus sarawakensis; R.K. Osborn |
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Immanus sarawakensis; R.K. Osborn |
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Immanus sarawakensis; R.K. Osborn |
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Immanus sarawakensis; R.K. Osborn |
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Immanus desectus; R.K. Osborn |
Taxonomy
Immanus Hulcr and Cognato, 2013: 100.
Diagnosis
This is the largest xyleborine genus with species ranging between 5.00−8.80 mm (Hulcr and Cognato 2013Hulcr and Cognato 2013:
Hulcr J, Cognato AI. 2013. Xyleborini of New Guinea: a taxonomic monograph. Thomas Say Publications in Entomology, Entomological Society of America, Lanham, Maryland, 176 pp.). Immanus can be distinguished by its large size; robust form (1.97−2.50 times as long as wide); truncatetruncate:
appearing cut off or suddenly shortened
declivitydeclivity:
downward slope of either the pronotum or elytra
; pronotumpronotum:
the dorsal surface of the thorax
anterioranterior:
the front or forward; opposite of posterior
margin with elevated carinacarina:
an elevated ridge or keel, not necessarily high or acute
or a row of 4−6 coarse asperities; pronotalpronotal:
pertaining to the pronotum
discdisc:
the flat central upper surface of any body part (e.g. pronotum and elytra) asperateasperate:
covered in asperities
; tibial denticlesdenticle:
a small tooth, the sides of which are equal and the tip is above the middle of the base reduced or absent on meso- and metatibiae; tibial edge very uneven and rugged; scutellumscutellum:
a shield-like sclerotized plate located at the midpoint of the elytral base
flat, flush with elytra; procoxae contiguous; and mycangial tufts absent.
May be confused with
Distribution
Paleotropical
Gallery system
The gallery system is branched and lies in one transverse plane (Kalshoven 1959bKalshoven 1959b:
Kalshoven LGE. 1959b. Studies on the biology of Indonesian Scolytoidea 4. Data on the habits of Scolytidae. Second part. Tijdschrift voor Entomologie 102: 135-173 + pls. 15-22.).
Remarks
The genus was recently reviewed by Beaver et al. 2019Beaver et al. 2019:
Beaver RA, Sittichaya W, Liu L-Y. 2019. A review of the genus Immanus Hulcr amp; Cognato with a key to species (Coleoptera: Curculionidae: Scolytinae: Xyleborini). Zootaxa 4585: 378-386. http://dx.doi.org/10.11646/zootaxa.4585.2.11 and one additional species has since been described (Wang et al. 2020Wang et al. 2020:
Wang J, Smith SM, and Cognato AI. 2020. Immanus virago , a new species from Borneo (Coleoptera, Curculionidae: Scolytinae: Xyleborini). The Coleopterists Bulletin 74: 438-440. https://doi.org/10.1649/0010-065X-74.2.438).
