Cyclorhipidion nemesis


  Cyclorhipidion nemesis  paratype lateral; S.M. Smith

Cyclorhipidion nemesis paratype lateral; S.M. Smith


  Cyclorhipidion nemesis  paratype dorsal; S.M. Smith

Cyclorhipidion nemesis paratype dorsal; S.M. Smith


  Cyclorhipidion nemesis  paratype declivity; S.M. Smith

Cyclorhipidion nemesis paratype declivity; S.M. Smith


  Cyclorhipidion nemesis  paratype frontal; S.M. Smith

Cyclorhipidion nemesis paratype frontal; S.M. Smith


Taxonomic history

Cyclorhipidion nemesis Smith & Cognato, 2022: 9.

Diagnosis

2.0−2.2 mm long (mean = 2.08 mm; n = 23); 2.75−3.07 times as long as wide. This species is distinguished by the short, steep declivitydeclivity:
downward slope of either the pronotum or elytra
that is approximately 25% of total elytralelytral:
pertaining to the elytra
length, armed with large tuberclestubercle:
a small knob-like or rounded protuberance of the exoskeleton
on interstriaeinterstria:
longitudinal spaces along the elytra between the striae, which is not as<br /> impressed and bear smaller punctures.
1 and 3, interstriaeinterstria:
longitudinal spaces along the elytra between the striae, which is not as<br /> impressed and bear smaller punctures.
2 never granulategranulate:
pertaining to a coarse, grainy surface texture
, posterolateralposterolateral:
relating to end of the side part/portion
margins rounded, declivitaldeclivital:
pertaining to the elytral declivity
interstriae 1 setaeseta:
small hair-like or scale-like structure
in one row, interstriaeinterstria:
longitudinal spaces along the elytra between the striae, which is not as<br /> impressed and bear smaller punctures.
2 setaeseta:
small hair-like or scale-like structure
uniseriate and small size (Smith and Cognato 2022Smith and Cognato 2022:
Smith SM, Cognato AI. 2022. New exotic beetles found in the United States among pseudocryptic Cyclorhipidion species (Coleoptera: Curculionidae: Scolytinae: Xyleborini) revealed via a multigene phylogeny. Insect Systematics and Diversity Insect Systematics and Diversity 6(4), 2. https://doi.org/10.1093/isd/ixac014
).

May be confused with

This species most closely resembles C. bodoanum, C. californicum, C. cognatoi Lin and Smith, 2022, and C. xeniolum.

This species is a part of the challenging C. pelliculosum species group. Species in the group are most easily distinguished by the rows of setaeseta:
small hair-like or scale-like structure
along declivitaldeclivital:
pertaining to the elytral declivity
interstriae 1and 2. See table below:

species

declivital declivital:
pertaining to the elytral declivity
interstriae 1 setaeseta:
small hair-like or scale-like structure
rows

declivital declivital:
pertaining to the elytral declivity
interstriae 2 setaeseta:
small hair-like or scale-like structure
rows

total body length (mm)

total body length/width ratio

lateral lateral:
pertaining to the side
profile of elytralelytral:
pertaining to the elytra
declivity

interstriae 2 granules

interstriae 1 & 3 granules

C. achlys Smith, Beaver, and Cognato, 2022

3

1

3.4

3.09

steeply rounded

absent

present

C. beaveri Lin and Smith, 2022

1

1

1.68−1.85

2.91−3.05

steeply rounded

present

present

C. bodoanum

2

1

1.6−2.4

2.5−3.29

steeply rounded

absent

present

C. californicum

2

2

2.2

3.14

steeply rounded

absent

present

C. cognatoi Lin and Smith, 2022

2

1

2.95−3.28

2.82−2.93

steeply rounded

absent

present

C. distinguendum

3

2

2.5−3.11

2.6−3.0

steeply rounded

absent

present

C. inarmatum

2

1

2.75−3.0

2.75−3.0

steeply rounded

absent

present

C. nemesis

1

1

2−2.2

2.75−3.07

steeply rounded

absent

present

C. pelliculosum

3

1

3.2−3.5

2.67−3.0

steeply rounded

rare, on basalbase:
point or edge closest to the body; opposite of apex
1/4

present

C. taedulum Smith, Beaver, and Cognato, 2022

2

1

2.7

3

steeply rounded

absent

present

C. tenuigraphum

2

2

2.7−3.0

2.5−3.0

steeply rounded

Often on apicalapex:
point or edge furthest from the body; opposite of base
1/3

present

C. triste

2

1

2.92−3.47

2.78

steeply rounded

absent

absent

C. xeniolum

1

1

1.65−1.8

3.09−3.4

obliquely truncateobliquely truncate:
nearly truncate but rounded not flat in lateral view

absent

present

Distribution

China (Yunnan) and established in the United States (Florida, Georgia, South Carolina) (Smith and Cognato 2022Smith and Cognato 2022:
Smith SM, Cognato AI. 2022. New exotic beetles found in the United States among pseudocryptic Cyclorhipidion species (Coleoptera: Curculionidae: Scolytinae: Xyleborini) revealed via a multigene phylogeny. Insect Systematics and Diversity Insect Systematics and Diversity 6(4), 2. https://doi.org/10.1093/isd/ixac014
, Smith et al. 2022cSmith et al. 2022c:
Smith, SM, Beaver RA, Pham HT, Cognato AI. 2022c. New species and new records of Xyleborini from the Oriental region, Japan and Papua New Guinea (Coleoptera: Curculionidae: Scolytinae). Zootaxa. In press.
)

DNA data

Sequences available for COI and CAD.

COI: MN199575 (holotype); OM867007; OP208788

CAD: MN199592 (holotype); OM891126; OP219692