Subfamily: Faboideae.
Phylogenetic Number: 3.26.01.
Tribe: Hypocalypteae.
Species Studied - Species in Genus: 3 studied; 3 in genus.
Fruit: A legume; unilocular; 0.2–6.5 cm long; 0.4–1.4 cm wide; 0.25 cm thick; 2–9 times longer than wide; with deciduous androecial sheath; with deciduous corolla; with persistent calyx; with calyx shorter than fruit; without orifice formed by curving of fruit or fruit segments; straight to curved (slightly); not plicate; not twisted; symmetrical, or asymmetrical; linear, or oblong, or obovate; with both sutures nearly straight; not inflated, or inflated; compressed, or terete; with beak; short tapered at apex; aligned with longitudinal axis of fruit; tapered at base; oblique with longitudinal axis of fruit; with the apex and base uniform in texture; coriaceous, or chartaceous; seed chambers externally visible; seed chambers with the raised seed chambers torulose; margin constricted, or not constricted; margin slightly constricted along both margins; margin without sulcus; margin plain; wing(s) absent; substipitate; with all layers dehiscing; splitting along suture(s). Dehiscence of valves along both sutures; medial and up and down; passive. Replum invisible. Epicarp dull; monochrome; reddish or blackish brown or brown, or black, or yellow (pale); with surface texture uniform; glabrous; eglandular; without spines; not smooth; with elevated features; veined; reticulately veined; not tuberculate; not exfoliating; without cracks; without embedded tissue, much thicker than epicarp, running from base to apex. Mesocarp present; thin; surface not veined; 1-layered; without balsamic vesicles; without fibers; without reniform canals; solid; coriaceous. Endocarp present; visible; dull; opaque; monochrome; tan; smooth; without adhering pieces of testa; nonseptate; chartaceous; not exfoliating; remaining fused to mesocarp and epicarp; without wings; entire. Seed(s) 1–6; length parallel with fruit length; neither overlapping nor touching; in 1 series. Funiculus of 1 length only; triangular; straight. Aril present; fleshy; when fleshy cupshaped; entire; covering less than 1/2 of seed; white.
Seed: 3.4–8 mm long; 2–3 mm wide; 1.3–3 mm thick; not overgrown; not angular; asymmetrical; reniform (oblong), or ovate; compressed; with surface smooth; with visible radicle and cotyledon lobes (faintly), or without visible radicle and cotyledon lobes; without external groove between radicle and cotyledon lobes; without hilar sinus; without umbo on seed faces; without medial ridge on each face. Cuticle not exfoliating; not inflated; not wrinkled. Testa present; without pieces of adhering epicarp; not adhering to endocarp; free from endocarp; dull and glossy, or dull; not modified by a bloom; colored; monochrome, or mottled to streaked; with frequent mottles; with frequent streaks; black to brown (dark); with black overlay (brownish); glabrous; smooth; coriaceous. Pleurogram absent. Pseudopleurogram absent. Fracture lines absent. Rim absent. Raphe visible, or not visible; from hilum through lens to base of seed and terminating; not bifurcating; darker than testa; reddish brown, or black; flush. Hilum present; partially concealed; concealed by aril; with faboid split; with the lips of the faboid split the same color as the rest of the hilum; larger than punctiform; 1 mm long; with straight outline; oblong; between cotyledon and radicle lobe, or marginal according to radicle tip; recessed; not within corona, halo, or rim. Lens discernible, or not discernible; with margins curved; circular (outline difficult to see); not in groove of raphe; adjacent to hilum; 1 mm from hilum; mounded; dissimilar color from testa; darker than testa; reddish brown, or black; not within corona, halo, or rim. Endosperm present; thick; not pluglike and not resembling tip of radicle; covering entire embryo; adnate to embryo. Cotyledons smooth; both outer faces convex; both the same thickness; both more or less of equal length; not folded; margin entire 180 degrees from base of radicle; similar at apex; not concealing radicle; entire over radicle; without lobes; with the interface division terminating at base of radicle; without margins recessed; yellow; inner face flat; glabrous on inner face. Embryonic axis deflexed; oblique to length of seed; without a joint evident between the radicle and the cotyledons. Radicle differentiated from cotyledon; linear; deflexed and parallel to cotyledon width; centered between cotyledons; less than 1/2 length of cotyledons. Plumule rudimentary; glabrous.