Subfamily: Mimosoideae.
Phylogenetic Number: 2.5.01A.
Tribe: Ingeae.
Species Studied - Species in Genus: 2 studied; 4 in genus.
Fruit: A loment (or a loment segment); (4.5–)5–12.5 cm long (Barneby and Grimes, 1996); 0.75–1.9 cm wide (Barneby and Grimes, 1996); 0.2–0.4 cm thick; 2–9 times longer than wide; with deciduous androecial sheath; with deciduous corolla; with deciduous calyx; without orifice formed by curving of fruit or fruit segments; straight, or curved; not plicate; not twisted; symmetrical, or asymmetrical; oblong, or linear, or falcate; with both sutures nearly straight, or both sutures parallelly curved; not inflated; flattened; with beak, or without beak; straight; with solid beak the same color and texture as fruit; rounded at apex, or tapered at apex (rarely); aligned with longitudinal axis of fruit; tapered at base; aligned with longitudinal axis of fruit; with the apex and base uniform in texture; leathery, or ligneous; seed chambers externally visible; seed chambers with the raised seed chambers not torulose; margin not constricted; margin without sulcus; margin embellished; margin with thickened sutural areas; wing(s) absent; nonstipitate, or substipitate (with pseudostipe, Barneby and Grimes, 1996); with the stipe 1–5 mm long; indehiscent. Replum invisible. Loment an intact article; indehiscent; segments (articles) conspicuous; segments (articles) (5–)5.5–9(–10) mm long (Barneby and Grimes, 1996); segments (articles) widest across seed area; segments (articles) with apical 1 different shape than middle one(s); segments (articles) oblong, or D-shaped (apical segment). Epicarp dull; monochrome; dark brown; with surface texture uniform; pubescent and indurate; with hairs erect; with 1 type of pubescence; minutely puberulent; with pubescence brown; pubescence denser near sutures, sparser centrally; with simple hairs; pliable; with hair bases plain; eglandular; without spines; not smooth; with elevated features; veined; transversely veined relative to fruit length; not tuberculate; not exfoliating; without cracks; without embedded tissue, much thicker than epicarp, running from base to apex. Mesocarp present, or absent; thick; surface not veined; 1-layered; without balsamic vesicles; without fibers; without reniform canals; solid; chartaceous. Endocarp present; visible; dull, or glossy; opaque; monochrome; tan; smooth, or smooth and scurfy (sometimes in a small area under the seed); without adhering pieces of testa; septate; with septa thicker than paper, firm; with septa eglandular; coriaceous; not exfoliating; remaining fused to mesocarp and epicarp; remaining fused to epicarp; without wings; entire. Seed(s) 8–13 (in fruits prior to separation of loments, Barneby and Grimes, 1996); length parallel with fruit length; neither overlapping nor touching; in 1 series. Funiculus 1–2 mm long; of 1 length only; filiform; convoluted, or straight. Aril absent.
Seed: 7.3–10.5 mm long; 4–6 mm wide; 1.5–2 mm thick; not overgrown; not angular; symmetrical; narrowly obovate, or irregular; flattened; with surface smooth; without visible radicle and cotyledon lobes; without hilar sinus; without umbo on seed faces; without medial ridge on each face. Cuticle not exfoliating; not inflated; not wrinkled. Testa present; without pieces of adhering epicarp; not adhering to endocarp; free from endocarp; dull; not modified by a bloom; colored; monochrome; brown; glabrous; smooth; chartaceous. Pleurogram present; 100 %. Pseudopleurogram absent. Fracture lines absent. Rim absent. Wing(s) absent. Raphe not visible. Hilum present; fully concealed; concealed by funicular remnant; without faboid split (assumed); punctiform; apical at apex of radicle tip; flush; not within corona, halo, or rim. Lens discernible; 0.1–0.5 mm long; with margins curved; circular, or elliptic; not in groove of raphe; adjacent to hilum; 0.1–0.2 mm from hilum; recessed; same color as testa; brown; not within corona, halo, or rim. Endosperm present; thin; not pluglike and not resembling tip of radicle; covering entire embryo; adnate to testa. Cotyledons smooth; both outer faces convex; both the same thickness; both more or less of equal length; not folded; margin entire 180 degrees from base of radicle; similar at apex; completely concealing radicle, or partially concealing radicle; split over radicle; without lobes; with the interface division terminating at base of radicle; without margins recessed; tan; inner face flat; glabrous on inner face. Embryonic axis parallel; parallel to length of seed; without a joint evident between the radicle and the cotyledons. Radicle differentiated from cotyledon; bulbose; lobe tip straight; straight with embryonic axis; centered between cotyledons; less than 1/2 length of cotyledons. Plumule well developed; glabrous.