Subfamily: Caesalpinioideae.
Phylogenetic Number: 1.4.38.
Tribe: Detarieae.
Group: Detarium.
Species Studied - Species in Genus: 3 studied; 5 in genus.
Fruit: A legume; unilocular; 2.5–6.5 cm long; 1.5–3.5 cm wide; 0.3–0.5 cm thick; length less than twice as long as width to 2–9 times longer than wide; with deciduous androecial sheath; with deciduous corolla; with deciduous calyx; without orifice formed by curving of fruit or fruit segments; straight; not plicate; not twisted; symmetrical, or asymmetrical; elliptic, or oblong to obovate; with 1 straight and 1 curved suture; widest near middle or D-shaped; not inflated; flattened; without beak; tapered at apex; aligned with longitudinal axis of fruit; tapered at base to rounded at base; aligned with longitudinal axis of fruit; with the apex and base uniform in texture; coriaceous to ligneous; seed chambers externally visible, or invisible; margin not constricted; margin without sulcus; margin plain; wing(s) absent; substipitate to nonstipitate (nearly); with all layers dehiscing; splitting along suture(s). Dehiscence of valves along 1 suture, or both sutures (ventral). Replum invisible. Epicarp dull; monochrome; blackish brown; with surface texture uniform; glabrous; eglandular; without spines; not smooth; with elevated features, or recessed features; not veined; not tuberculate; glandular dotted; glandularly punctate; not exfoliating; without cracks, or with cracks (in G. kisantuense); cracking oblique to fruit length; without embedded tissue, much thicker than epicarp, running from base to apex. Mesocarp present; surface not veined; 2-layered; without balsamic vesicles; with fibers; without reniform canals; with fibers solid or over mealy tissue; ligneous to coriaceous. Endocarp present; visible; dull; opaque; monochrome; brown to reddish brown; without adhering pieces of testa; nonseptate; not exfoliating; remaining fused to mesocarp and epicarp; entire. Seed(s) 1–2; length oblique to fruit length to transverse to fruit length; neither overlapping nor touching; in 1 series. Funiculus 0.1–2 mm long; of 1 length only; thick; straight. Aril present (absent in G. kisantuense); dry; when dry shape not stated but the apex is indicated to have acute or obtuse lobes which may be erect or reflexed; entire; covering less than 1/2 of seed (degree of coverage not stated).
Seed: 11–23 mm long; 6.5–17 mm wide; 3–4 mm thick; not overgrown; not angular; symmetrical; oblong; compressed; with surface smooth; without visible radicle and cotyledon lobes; without hilar sinus; without umbo on seed faces; without medial ridge on each face. Cuticle not exfoliating; not inflated; not wrinkled. Testa present; without pieces of adhering epicarp; not adhering to endocarp; free from endocarp; dull; not modified by a bloom; colored; monochrome; dark reddish brown; glabrous; smooth; osseous. Pleurogram absent. Pseudopleurogram absent. Fracture lines present; transversely reticulate. Rim absent. Wing(s) absent. Raphe not visible. Hilum present; visible, or fully concealed; concealed by funicular remnant; without faboid split; larger than punctiform; 0.1–0.5 mm long; with curved outline; elliptic; subapical to radicle tip; recessed; not within corona, halo, or rim. Lens discernible; 0.1–0.5 mm long; with margins straight; linear; not in groove of raphe; mounded; similar color as testa; darker than testa; not within corona, halo, or rim. Endosperm absent. Cotyledons smooth; both outer faces convex; both the same thickness; both more or less of equal length; not folded; margin entire 180 degrees from base of radicle; similar at apex; completely concealing radicle; notched at radicle and split over radicle, or notched at radicle (only); with the interface division terminating at base of radicle; without margins recessed; inner face flat; glabrous on inner face. Embryonic axis straight; parallel to length of seed to oblique to length of seed; without a joint evident between the radicle and the cotyledons. Radicle differentiated from cotyledon; centered between cotyledons. Plumule rudimentary; glabrous.
Seed, cotyledon, embryo, and testa: G. kisantuense (F.M.C. Vermoesen) ex E.A.J. De Wildeman) J.J.G. Léonard - bottom far left seed topography, top left cotyledon notched and concealing radicle (L) and embryonic axis (R); G. pierreanum (H.A.T. Harms) J.J.G. Léonard - testa SEMs; G. spp. - bottom left center seeds.
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