Subfamily: Caesalpinioideae.
Phylogenetic Number: 1.4.68.
Tribe: Detarieae.
Group: Macrolobium.
Species Studied - Species in Genus: 8 studied; 26 in genus.
Fruit: A legume; unilocular; 10–30 cm long; 4–9 cm wide; 0.4–1.3 cm thick; 2–9 times longer than wide; with deciduous androecial sheath; with deciduous corolla; with deciduous calyx; without orifice formed by curving of fruit or fruit segments; straight to curved; not plicate; not twisted; asymmetrical; oblong to oblanceolate; not inflated; compressed; without beak; tapered at apex; oblique with longitudinal axis of fruit; rounded at base; right angled with longitudinal axis of fruit; with the apex and base uniform in texture; ligneous; seed chambers externally invisible; margin not constricted; margin without sulcus; margin plain; wing(s) absent; nonstipitate; with all layers dehiscing; splitting along suture(s). Dehiscence of valves along both sutures; active; with valves elastically and separately enrolling. Replum invisible. Epicarp dull; monochrome; reddish brown; with surface texture uniform; densely reddish velvety brown pubescent and indurate; with hairs erect; with simple hairs; eglandular; without spines; not smooth; veined; longitudinally veined relative to fruit length, obliquely veined relative to fruit length, and reticulately veined (either 1–3 prominent longitudinal veins with ot without 1 major branch and with obliquely reticulate veins or 1 longitudinal vein near ventral margin giving rise to numerous oblique reticulate veins); not tuberculate; exfoliating; without cracks; without embedded tissue, much thicker than epicarp, running from base to apex. Mesocarp present; surface not veined; 1-layered; without balsamic vesicles; without fibers; without reniform canals; solid; ligneous. Endocarp present; visible; dull; opaque; monochrome; brown; without adhering pieces of testa; nonseptate; not exfoliating; remaining fused to mesocarp and epicarp; entire. Seed(s) 1–8; length transverse to fruit length; neither overlapping nor touching; in 1 series. Funiculus 0.1–0.19 mm long; of 1 length only; thick; nearly straight. Aril absent.
Seed: 20–50 mm long; 15–50 mm wide; 5–11 mm thick; not overgrown; not angular; symmetrical; nearly circular to oblong; compressed; with surface smooth; without visible radicle and cotyledon lobes; without hilar sinus; without umbo on seed faces; without medial ridge on each face. Cuticle not exfoliating; not inflated; not wrinkled. Testa present; without pieces of adhering epicarp; not adhering to endocarp; free from endocarp; glossy; not modified by a bloom; colored; monochrome; reddish to blackish brown; glabrous; not smooth; with elevated features; rugose; coriaceous. Pleurogram absent. Pseudopleurogram absent. Fracture lines absent. Rim absent. Wing(s) absent. Raphe not visible. Hilum present; fully concealed; concealed by funicular remnant; without faboid split; larger than punctiform; 0.1–4 mm long; with curved outline; elliptic; apical at apex of radicle tip; recessed; not within corona, halo, or rim. Lens not discernible. Endosperm absent. Cotyledons smooth; both outer faces convex; both the same thickness; both more or less of equal length; not folded; margin entire 180 degrees from base of radicle; similar at apex; partially concealing radicle (only tip exposed); notched at radicle and split over radicle; with the interface division terminating at base of radicle; without margins recessed; inner face flat; glabrous on inner face. Embryonic axis straight; parallel to length of seed to perpendicular to length of seed; without a joint evident between the radicle and the cotyledons. Radicle differentiated from cotyledon; centered between cotyledons. Plumule well developed; glabrous.
Seed, cotyledon, embryo, and testa: G. dewevrei (E.A.J. De Wildeman) J.J.G. Léonard - top left center seed topography, top far left cotyledon notched and split and concealing all but tip of radicle (above) and embryonic axis (below); G. dinklagei H.A.T. Harms - testa SEMs; G. limba (A.C. Gilliot) J.J.G. Léonard - bottom left center seed; G. sp. - bottom far left seed topography.
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