Subfamily: Faboideae.
Phylogenetic Number: 3.10.27.
Tribe: Phaseoleae.
Subtribe: Diocleinae.
Species Studied - Species in Genus: 18 studied; 50 in genus.
Fruit: A legume; unilocular; 2.5–8 cm long; 0.4–1.2 cm wide; 0.1–0.3 cm thick; 2–9 times longer than wide; with persistent androecial sheath (with or without the keel), or deciduous androecial sheath; with persistent corolla (with or without androecial sheath), or deciduous corolla; with keel petal; with persistent calyx, or deciduous calyx; with calyx shorter than fruit; without orifice formed by curving of fruit or fruit segments; straight, or curved (slightly); not plicate; not twisted; asymmetrical, or symmetrical; linear, or falcate; with both sutures parallelly curved; not inflated; compressed, or flattened; without beak; tapered at apex, or short tapered at apex; aligned with longitudinal axis of fruit, or oblique with longitudinal axis of fruit; tapered at base; aligned with longitudinal axis of fruit; with the apex and base uniform in texture; chartaceous to coriaceous; seed chambers externally invisible, or visible; seed chambers with the raised seed chambers not torulose; margin not constricted; margin without sulcus; margin plain, or embellished; margin with thickened sutural areas (slightly); wing(s) absent; nonstipitate; with all layers dehiscing; splitting along suture(s). Dehiscence of valves along both sutures; apical and down; active; with valves twisting. Replum invisible. Epicarp dull; monochrome, or multicolored; mottled; brown; with brown overlay (darker); mottling color combination constant; with mottling over seed chambers; with surface texture uniform; pubescent and indurate, or pubescent but soon deciduous; with hairs erect; with 1 type of pubescence, or 2 types of pubescence; densely tomentose, or velutinous, or sericeous, or tomentose and villous; with pubescence white, or golden; with pubescence uniformly distributed; with simple hairs; pliable; with hair bases plain; eglandular; without spines; smooth; not veined; not tuberculate; not exfoliating; without cracks; without embedded tissue, much thicker than epicarp, running from base to apex. Mesocarp present; thin; surface not veined; 1-layered; without balsamic vesicles; without fibers; without reniform canals; solid; coriaceous. Endocarp present; visible; dull; opaque; monochrome; brown to tan; cracked; without adhering pieces of testa; subseptate; with septa thin (tissue paper-like), flexible; with septa eglandular; chartaceous; exfoliating; separating from mesocarp; without wings; entire. Seed(s) 5–10; length parallel with fruit length, or oblique to fruit length; neither overlapping nor touching; in 1 series. Funiculus 0.5–4 mm long; of 1 length only; filiform; curved, or triangular. Aril present; dry; when dry 2-lipped rim-aril; entire; covering less than 1/2 of seed; without tongue (or flap) on lips of 2-lipped rim-aril, or with tongues (or flap) on lips of 2-lipped rim-aril; with 2 tongues or flaps, 1 on each lip of 2-lipped rim-aril; tan.
Seed: 3.5–7.8 mm long; 2.6–4.5 mm wide; 1.7–3.7 mm thick; not overgrown; not angular; symmetrical, or asymmetrical; D-shaped, or elliptic, or ovate, or reniform; terete, or compressed; with surface smooth; without visible radicle and cotyledon lobes; without hilar sinus; without umbo on seed faces; without medial ridge on each face. Cuticle not exfoliating; not inflated; not wrinkled. Testa present; without pieces of adhering epicarp; not adhering to endocarp; free from endocarp; dull; not modified by a bloom; colored; monochrome, or mottled; with frequent mottles, or infrequent mottles; brown, or orange (brownish), or tan; with brown overlay, or tan overlay; glabrous; smooth; chartaceous. Pleurogram absent. Pseudopleurogram absent. Fracture lines absent. Rim absent. Wing(s) absent. Raphe not visible. Hilum present; visible; with faboid split; with the lips of the faboid split the same color as the rest of the hilum; larger than punctiform; 1–4.5 mm long; with curved outline, or straight outline; elliptic; linear; apical according to radicle tip but marginal according to seed length; raised; within rim; rim color of testa, or darker than testa. Lens discernible; 0.5–0.7 mm long; with margins straight; linear; not in groove of raphe; confluent with hilum; flush; similar color as testa; darker than testa; brown, or red; within halo; halo color of testa, or darker than testa. Endosperm present; trace; not pluglike and not resembling tip of radicle; restricted to region of embryo; adnate to testa. Cotyledons smooth; both outer faces convex; both the same thickness; both more or less of equal length; not folded; margin entire 180 degrees from base of radicle, or not entire 180 degrees from base of radicle; slightly notched; similar at apex; partially concealing radicle; split over radicle; without lobes; with the interface division terminating at base of radicle; without margins recessed; white; inner face flat; glabrous on inner face. Embryonic axis right angled; oblique to length of seed, or perpendicular to length of seed; without a joint evident between the radicle and the cotyledons. Radicle differentiated from cotyledon; linear, or triangular; lobe tip curved; oblique to cotyledons; centered between cotyledons; less than 1/2 length of cotyledons. Plumule well developed; glabrous.