Subfamily: Mimosoideae.
Phylogenetic Number: 2.5.19.
Tribe: Ingeae.
Species Studied - Species in Genus: 13 studied; 94 in genus.
Fruit: A legume; unilocular; 3–50 cm long; 1–8.5 cm wide; 0.4–4 cm thick; 2–9 times longer than wide; with deciduous androecial sheath; with deciduous corolla; with deciduous calyx; without orifice formed by curving of fruit or fruit segments; straight to 0.5-coiled to 1-coiled to 1.5-coiled to 2-coiled (spirally); not plicate; twisted, or not twisted; symmetrical, or asymmetrical; falcate, or C-shaped, or reniform, or irregular; with both sutures parallelly curved, or both sutures unequally curved; not inflated; flattened to terete; without beak; rounded at apex; oblique with longitudinal axis of fruit, or aligned with longitudinal axis of fruit; tapered at base; oblique with longitudinal axis of fruit; with the apex and base uniform in texture; when fresh fleshy and coriaceous, or ligneous (when dry); seed chambers externally visible, or invisible; margin not constricted to constricted; margin constricted only on 1 margin to slightly constricted only on 1 margin; margin without sulcus; margin plain; wing(s) absent; stipitate, or substipitate; with the stipe up to 40 mm long; tardily with all layers dehiscing; splitting along suture(s). Dehiscence of valves along 1 suture (along dorsal or occasionally ventral margin); medial and up and down; active; with valves twisting (remaining attached to sutures). Replum invisible. Epicarp dull; monochrome; various shades of brown to red, or green; with surface texture uniform; glabrous to pubescent and indurate; with hairs erect; with simple hairs; eglandular; without spines; smooth to not smooth; with elevated features; veined; reticulately veined; not tuberculate; rugose; not exfoliating; without cracks; without embedded tissue, much thicker than epicarp, running from base to apex. Mesocarp absent, or present; surface not veined; 1-layered; without balsamic vesicles; without reniform canals; fibrous throughout; ligneous. Endocarp present; visible; dull to glossy; opaque; monochrome; orange to red, or brown to gray, or white; rugose; without adhering pieces of testa; subseptate to nonseptate; not exfoliating; remaining fused to mesocarp and epicarp; entire. Seed(s) 2–15; length transverse to fruit length; neither overlapping nor touching; in 1 series. Funiculus 1–15 mm long; of 1 length only; thick to filiform; straight to hooked, or plicate. Aril absent.
Seed: 8–45 mm long; 7–23 mm wide; 4–30 mm thick; not overgrown; not angular; symmetrical; circular to elliptic, or oblong to reniform; terete to compressed; with surface grooved (1 on each face); longitudinal; without visible radicle and cotyledon lobes; without hilar sinus; without umbo on seed faces; without medial ridge on each face. Cuticle not exfoliating; not inflated; not wrinkled. Testa present; without pieces of adhering epicarp; not adhering to endocarp; free from endocarp; glossy, or glaucous (occasionally); not modified by a bloom; colored; monochrome; bluish black to yellow, or brown (occasionally with bloom); glabrous; not smooth to smooth; with elevated features, or recessed features; rugose, or wrinkled; chartaceous to osseous. Pleurogram absent. Pseudopleurogram absent. Fracture lines absent. Rim absent. Wing(s) absent. Raphe not visible. Hilum present; fully concealed; concealed by funicular remnant (orange-brown to reddish-brown in color); without faboid split; punctiform, or larger than punctiform; up to 5 mm long; with curved outline; circular, or elliptic; apical at apex of radicle tip; flush to raised; not within corona, halo, or rim. Lens not discernible. Endosperm absent. Cotyledons not smooth (rugose), or smooth; both outer faces flat; both the same thickness; both more or less of equal length; not folded; margin entire 180 degrees from base of radicle; similar at apex; completely concealing radicle; split over radicle; with the interface division terminating at base of radicle; without margins recessed; inner face flat; glabrous on inner face. Embryonic axis straight; parallel to length of seed; without a joint evident between the radicle and the cotyledons. Radicle differentiated from cotyledon; straight with embryonic axis; centered between cotyledons. Plumule well developed; glabrous.
Seed, cotyledon, and embryo: A. aruense (O. Warburg) H.C.D. de Wit - 1st row cotyledons concealing radicle (L) and embryonic axis (R); A. ellipticum (Blume) I.C. Nielsen - 2nd row top seed topography, 3rd row bottom seed; A. jiringa (Jack) I.C. Nielsen - 4th row seed; A. lucyi F.J.H. von Mueller - 3rd row top seed topography, 2nd row bottom seed.
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