Native
Exotic, but established
The tribe Olethreutini contains over 1,100 described species, making it the second-most diverse tribe of Olethreutinae, behind only Eucosmini. Several subgroups may deserve tribal status, but future work is needed. Horak (2006)Horak (2006):
Horak, M. 2006. Olethreutine moths of Australia (Lepidoptera: Tortricidae). Monographs on Australian Lepidoptera, Vol. 10. 522 pp. included the former tribe Gatesclarkeanini as the Gatesclarkeana group within the Olethreutini. Regier et al. (2012) synonymized the former tribes Bactrini and Endothiini under Olethreutini as well. Its members occur worldwide, although the greatest diversity is in southeast Asia and the Holarctic. There are no synapomorphies that define the entire tribe, although a hindwing with veins M3 and CuA1 separate and a sacculussacculus:
the ventral margin of the male valva
with two clusters of spines unite some species.
Many species can be recognized as members of Olethreutini by the short, upturned labial palpilabial palpi:
a pair of sensory appendages that project from the lower part of the head; usually covered in scales and three-segmented
; and relatively broad forewing with a complex and irregular pattern of fasciaefasciae:
plural of "fascia"
. However, many exceptions exist. For example, Bactra have moderate or long, porrectporrect:
extending forward horizontally
labial palpilabial palpi:
a pair of sensory appendages that project from the lower part of the head; usually covered in scales and three-segmented
; a narrow forewing and often relatively simple forewing markings.
Most Olethreutini tend to be highly polyphagous leafrollers on various trees and shrubs, but exceptions exist here as well. Bactra are stem-borers on grasses (Poaceae), sedges (Cyperaceae) and rushes (Juncaceae), whereas some species of Olethreutes may be monophogous. Only a select few species of Olethreutini are considered serious pests (e.g., Lobesia botrana, Paralobesia viteana). We include these few pest species as well as several common or similar-looking non-pest species on this site.
The most recent comprehensive guide to the Nearctic Olethreutini is Heinrich (1926)Heinrich (1926):
Heinrich, C. 1926. Revision of the North American moths of the subfamilies Laspeyresiinae and Olethreutinae. Bulletin of the United States National Museum. 132: 1-216., but more recent regional (Miller 1987; Gilligan 1987) and genus-level (Heppner 1994Heppner 1994:
Heppner, J. B. 1994. Episimus moths of North America (Lepidoptera: Tortricidae). Holarctic Lepidoptera. 1: 83-107.; Razowski & Brown 2008; Royals et al. 2019Royals et al. 2019:
Royals, H. R., J-F. Landry, T. M. Gilligan. 2019. Paralobesia (Lepidoptera: Tortricidae), a systematic revision. Memoirs of the Lepidopterists#39; Society, No. 6. Washington, D.C. 149 pp.) treatments exist. There are major discrepancies between how North American and European workers treat certain genera of Olethreutini. See the Olethreutes spp. page for more information.
Included species:
Exotic, but established species
Lobesia botrana
Native species
Bactra spp.
Endothenia hebesana
Episimus argutana
Eumarozia malachitana
Hedya chionosema
Hedya separatana
Olethreutes spp.
Paralobesia spp.
Paralobesia viteana
Phiaris siderana
View full screen host table here