Exotic
Common names: chestnut leafroller
Synonyms: herrichiana (Carpocapsa), juliana (Philalcea), nimbana (Carpocapsa), trinotana (Tortrix)
FWL: 6.5-8.5 mm
Forewings are white and gray with a large ocellusocellus:
forewing pattern element - an ovoid region anterior to the tornus; adult head - a simple insect "eye" located dorsal to the compound eye 
consisting of a leaden-metallic center bar surrounded by black and yellow dashes. Adults are not sexually dimorphic. Males lack a forewing costal foldforewing costal fold:
a flap or fold at the base of the forewing that contains specialized sex scales
. Hindwings are brown.
Male genitalia are characterized by the absence of an uncusuncus:
a sclerotized process which is fused to the posterodorsal margin of tergum IX
and several cornuticornuti:
spines used to anchor the male vesica in the female bursa during copulation 
in the vescia. Female genitalia are characterized by two thornlike signasigna:
plural of "signum"
and a short ductus bursaeductus bursae:
a membranous tube connecting the ostium bursae to the corpus bursae
with a sclerotizedsclerotized:
hardened; usually in reference to larval structures or adult genitalia 
ring.
The following account is summarized from Brown & Komai (2008).
Mature larva approximately 13-15 mm in length; head light brown or yellowish brown with darker mottling; prothoracic shieldprothoracic shield:
a sclerotized plate on the dorsal surface of the prothorax 
whitish with dark dots; body grayish white with large, black, conspicuous pinaculapinacula:
flattened sclerotized plates on a caterpillar that bear the setae
; anal shieldanal shield:
a sclerotized plate on the dorsal surface of the last abdominal segment (in larvae) 
brown with darker brown dots; anal fork present; SV group on A1, 2, 7, 8, 9 most often 3:3:2:2:1 but variable.
The majority of Pammene are Palearctic, with over 90 species present in the region. Only six species are present in the Nearctic and all can be distinguished from P. fasciana by wing pattern. A genitalic dissection can be used to confirm the identity of questionable or worn individuals.
Other tortricid Castanea pests include Cydia fagiglandana, Cydia glandicolana, Cydia kurokoi, Cydia splendana, and Fibuloides aestuosa. Brown and Komai (2008) provide a description and key to larvae of these species. Pammene fasciana larvae can be distinguished by the following characters: whitish abdomen with large, dark, conspicuous pinaculapinacula:
flattened sclerotized plates on a caterpillar that bear the setae; SV setal counts on A1,2,7,8,9 as 3:3:2:2(1):1; three L setae on A9 on single pinaculumpinaculum:
singular of "pinacula"
; distance between V setae on A9 greater than distance between Vs on A8; crochetscrochets:
hooked spines on the prolegs of lepidopterous larvae 
on prolegs uniordinal (or irregularly biordinal); number of crochetscrochets:
hooked spines on the prolegs of lepidopterous larvae on prolegs 30-33; anal combanal comb:
a toothed structure on the last abdominal segment used to eject frass away from the feeding larva; also termed "anal fork" 
present.
The following account is summarized from Bogenschütz (1991).
Pammene fasciana completes a single generation per year. Adults are present June-July. Males fly after sunrise and again in the afternoon; females also fly at night.
Females lay eggs singly or in small groups along the veins on the leaves of the host plant. Larvae bore into nuts and feed inside. When the nuts fall from the tree, the larvae bore out of the nut and create a cocoon under tree bark or moss in which to overwinter. Pupation occurs the following spring.
Larvae of Pammene fasciana are important pests of chestnut (Castanea sativa). Other preferred hosts include Quercus and Fagus.
| Host plant | Host plant family | Reference(s) |
| Castanea sativa | Fagaceae | Bradley et al. 1979Bradley et al. 1979: Bradley, J. D., Tremewan, W. G., Smith, A. 1979. British Tortricoid Moths, Tortricidae: Olethreutinae. The Ray Society, London. 336 pp.; Rotundo et al. 1984Rotundo et al. 1984: Rotundo, G., Giacometti, R., Rotundo, A. 1984. Individuazione degli attrativi sessuali dei torticidi delle castagne mediante studio elettroantennografico e di campo. Bollettino Laboratori Entomologia Agraria Filippo Silvestri. 41: 211-224 [In Italian].; Rotundo & Tremblay 1993; Mansilla & Salinero 1993 |
| Castanea sp. | Fagaceae | USDA/APHIS interceptionsUSDA/APHIS interceptions: USDA/APHIS interceptions. Based on larvae intercepted by USDA-APHIS personnel at U.S. ports-of-entry and identified by Systematic Entomology Laboratory personnel.; MacKay 1959MacKay 1959: MacKay, M. R. 1959. Larvae of the North American Olethreutidae (Lepidoptera). Canadian Entomologist, Supplement 10: 1-338.; Rotundo & Giacometti 1986; Martin et al. 1998Martin et al. 1998: Martin, A., Baronio, P., Baldassari, N., Rocchetta, G. 1998. I lepidotteri torticidi del castagno ( Pammene fasciana (L.), Cydia fagiglandana (Zel.) e Cydia splendana (Hb.)) valutati come un#39;unica entita di danno. Bolletim Istituto Entomol. quot;Guido Grandiquot; Univ. Bologna. 52: 105-114. |
| Fagus sp. | Fagaceae | Disque 1908Disque 1908: Disque, H. 1908. Versuch einer microlepidopterologischen Botanik. Deutsch Entomologische Zeitschrift Iris. 21: 34-147. |
| Fagus sylvatica | Fagaceae | Bradley et al. 1979Bradley et al. 1979: Bradley, J. D., Tremewan, W. G., Smith, A. 1979. British Tortricoid Moths, Tortricidae: Olethreutinae. The Ray Society, London. 336 pp. |
| Quercus robur | Fagaceae | Disque 1908Disque 1908: Disque, H. 1908. Versuch einer microlepidopterologischen Botanik. Deutsch Entomologische Zeitschrift Iris. 21: 34-147. |
| Quercus suber | Fagaceae | Soria & Ocete 1996 |
| Quercus sp. | Fagaceae | Bradley et al. 1979Bradley et al. 1979: Bradley, J. D., Tremewan, W. G., Smith, A. 1979. British Tortricoid Moths, Tortricidae: Olethreutinae. The Ray Society, London. 336 pp. |
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Pammene fasciana is distributed from western Europe east to Ukraine.
