Uribea

Taxonomy

Uribea A. Dugand & R.C. Romero-Castañeda In Dugand, Mutisia 27: 5. 1 Sep 1962.

Subfamily: Faboideae.
Phylogenetic Number: 3.2.04.
Tribe: Sophoreae.
Group: Myroxylon.
Species Studied - Species in Genus: 1 studied; 1 in genus.

Description

Fruit: A legumelegume:
usually dry, dehiscent fruit derived from a single carpel that opens along two longitudinal sutures
; unilocular; 3–7 cm long; 2–2.7 cm wide; 1.4–1.8 cm thick; 2–9 times longer than wide; with deciduous androecial sheath; with deciduous corolla; with deciduous calyx; without orifice formed by curving of fruit or fruit segments; straight; not plicate; not twisted; asymmetrical; elliptic, or obovate, or obliquely obovate, or reniform; with both sutures parallelly curved, or both sutures unequally curved; somewhat inflated, or not inflated; terete; with beak; straight, or declined; with solid beak the same color and texture as fruit; rounded at apex; aligned with longitudinal axis of fruit, or oblique with longitudinal axis of fruit; rounded at base, or tapered at base, or short tapered at base; aligned with longitudinal axis of fruit; with the apex and base uniform in texture; coriaceous; seed chambers externally invisible; margin not constricted; margin without sulcus; margin plain; wing(s) absent; stipitate, or substipitate; with the stipe 2–10 mm long; indehiscent. Epicarp dull; monochrome; reddish brown or brown; with surface texture uniform; glabrous; eglandular; without spines; not smooth; with elevated features, or recessed features; not veined; not tuberculate; papillose; pitted; not exfoliating; with cracks, or without cracks; cracking transverse to fruit length; without embedded tissue, much thicker than epicarp, running from base to apex. Mesocarp present; thick; 1-layered; without balsamic vesicles; without fibers; without reniform canals; fleshy; fleshy. Endocarp present; concealed; concealed by fleshy mesocarp; without wings. Seed(s) 1–4; length transverse to fruit length; neither overlapping nor touching; in 1 series. Aril present; fleshy; when fleshy topknotlike; entire; covering less than 1/2 of seed; brown.

Seed: 14–19 mm long; 8–12 mm wide; 7–10 mm thick; not overgrown; angular, or not angular; asymmetrical; oblong, or obovate; quadrangular, or compressed; with surface smooth; without visible radicle and cotyledon lobes; without hilar sinus; without umbo on seed faces; without medial ridge on each face. Cuticle not exfoliating; not inflated; not wrinkled. Testa present; with pieces of adhering epicarp; not adhering to endocarp; free from endocarp; dull; not modified by a bloom; colored; monochrome; black to brown (dark); glabrous; smooth and not smooth; with elevated features; bearing endocarp remnants; chartaceous. Pleurogram absent. Pseudopleurogram absent. Fracture lines absent. Rim absent. Wing(s) absent. Raphe not visible. Hilum present; fully concealed; concealed by aril; larger than punctiform; 3–3.5 mm long; with curved outline; elliptic; subapical to radicle tip; recessed; not within corona, halo, or rim. Lens not discernible. Endosperm present; thick; not pluglike and not resembling tip of radicle; covering entire embryo; adnate to testa. Cotyledons smooth, or not smooth; convoluted; outer face of one cotyledon concave and other cotyledon convex; both the same thickness; both more or less of equal length; not folded; margin entire 180 degrees from base of radicle; similar at apex; partially concealing radicle; notched at radicle; with lobes; with lobes overlapping; with basal groin formed by lobes; with the interface division terminating at base of radicle; without margins recessed; brown; inner face wavy; glabrous on inner face. Embryonic axis oblique; oblique to length of seed; without a joint evident between the radicle and the cotyledons. Radicle differentiated from cotyledon; bulbose; lobe tip curved; oblique to cotyledons; centered between cotyledons; less than 1/2 length of cotyledons. Plumule rudimentary; glabrous.

Distribution

Costa Rica and Colombia.

New World; Costa Rica in Central America and South America (Colombia).

Tribal Notes

Tribe Sophoreae

Polhill (1981b) stated that the Sophoreae s.l. is a tribe of convenience between the Caesalpinioideae and the bulk of the Papilionoideae, sharply defined from neither. He transferred four genera from Sophoreae into the Swartzieae (1), Amburana (3.1.15), Ateleia (3.1.13), Cyathostegia (3.1.14), and Holocalyx (3.1.12), following Herendeen's (1995) cladistic analyses. Herendeen performed cladistic analyses for all Swartzieae genera, sensu Cowan (1981), 19 Sophoreae genera, and three Caesalpinioideae genera. He concluded that Swartzieae is polyphyletic and that it should be disbanded and its genera transferred to Sophoreae. Preliminary rbcL data (Doyle et al. 1997) supported his conclusions.