Sympetalandra

Taxonomy

Sympetalandra O. Stapf Hooker's Icon. Pl. 2721. Sep 1891.

Subfamily: Caesalpinioideae.
Phylogenetic Number: 1.1.40.
Tribe: Caesalpinieae.
Group: Dimorphandra.
Species Studied - Species in Genus: 3 studied; 5 in genus.

Description

Fruit: A legumelegume:
usually dry, dehiscent fruit derived from a single carpel that opens along two longitudinal sutures
; unilocular; 16–30 cm long; 1.5–5 cm wide; 0.4–2.5 cm thick; 2–9 times longer than wide to more than 9 times longer than wide; with deciduous androecial sheath; with deciduous corolla; with deciduous calyx; without orifice formed by curving of fruit or fruit segments; straight to curved; not plicate; not twisted; asymmetrical; oblong to linear; not inflated; compressed to terete; without beak; rounded at apex to tapered at apex; aligned with longitudinal axis of fruit; long tapered at base; aligned with longitudinal axis of fruit; with the apex and base uniform in texture; ligneous; seed chambers externally visible; margin constricted; margin constricted along both margins (in 2-seeded fruits), or slightly constricted along both margins (in 1-seeded fruits); margin without sulcus; margin plain; wing(s) absent; stipitate; with the stipe 6–20 mm long; indehiscent (remaining attached to sutures). Replum invisible. Epicarp dull; monochrome; reddish brown; with surface texture uniform; glabrous; eglandular; without spines; not smooth; with elevated features; veined; longitudinally veined relative to fruit length; not tuberculate; not exfoliating; without cracks; without embedded tissue, much thicker than epicarp, running from base to apex. Mesocarp present; surface not veined; 2-layered; without balsamic vesicles; with fibers; without reniform canals; with fibers over solid layer; ligneous. Endocarp present; visible; almost glossy; opaque; monochrome; reddish brown; smooth; without adhering pieces of testa; septate; not exfoliating; remaining fused to mesocarp and epicarp; entire. Seed(s) 1–2; length parallel with fruit length; neither overlapping nor touching; in 1 series. Funiculus 0.1–0.9 mm long; of 1 length only; thick; straight. Aril absent.

Seed: 40 mm long; 30 mm wide; 15 mm thick; not overgrown; not angular; symmetrical; oblong; compressed; with surface smooth; without visible radicle and cotyledon lobes; without hilar sinus; without umbo on seed faces; without medial ridge on each face. Cuticle not exfoliating; not inflated; not wrinkled. Testa present; without pieces of adhering epicarp; not adhering to endocarp; free from endocarp; dull; not modified by a bloom; colored; monochrome; reddish brown; glabrous; not smooth; with elevated features; rugose; osseous. Pleurogram absent. Pseudopleurogram absent. Fracture lines absent. Rim absent. Wing(s) absent. Raphe visible (from hilum across base to 180 degrees from hilum); not bifurcating; color of testa. Hilum present; fully concealed; concealed by funicular remnant; without faboid split; larger than punctiform; 0.1–5 mm long; with curved outline; elliptic; subapical to radicle tip (nearly marginal); raised; within halo; halo darker than testa. Lens not discernible. Endosperm absent. Cotyledons smooth; both outer faces convex; both the same thickness; both more or less of equal length; with only 1 folded; sufficiently folded for inner face to touch itself; portions of inner folded face unequal; margin not entire 180 degrees from base of radicle; similar at apex; partially concealing radicle; with lobes; with lobes overlapping; with basal groin formed by lobes; with the interface division terminating at base of radicle; without margins recessed; inner face flat; glabrous on inner face. Embryonic axis straight; slightly oblique to length of seed; without a joint evident between the radicle and the cotyledons. Radicle differentiated from cotyledon; centered between cotyledons. Plumule rudimentary; glabrous.

Distribution

Malaya, Borneo, Philippine Islands, and Lesser Sunda Islands.

Old World; Indonesia and the Philippines (Malaysia and Philippine Islands).

Generic Notes

Van Steenis (1975) revised Sympetalandra.

 Fruit and seed:  S. borneensis  O. Stapf - top 1-seeded fruit, bottom 2-seeded fruit with seed in situ.
Fruit and seed: S. borneensis O. Stapf - top 1-seeded fruit, bottom 2-seeded fruit with seed in situ.
 Seed, cotyledon, embryo, and testa:  S. borneensis  O. Stapf - far left cotyledon split and groined and concealing radicle (above) and embryonic axis (below), top left center seed topography, bottom left center seed, testa SEMs.
Seed, cotyledon, embryo, and testa: S. borneensis O. Stapf - far left cotyledon split and groined and concealing radicle (above) and embryonic axis (below), top left center seed topography, bottom left center seed, testa SEMs.