Pseudosindora

Taxonomy

Pseudosindora C.F. Symington Proc. Linn. Soc. London 155: 285. 9 Jun 1944.

Subfamily: Caesalpinioideae.
Phylogenetic Number: 1.4.45.
Tribe: Detarieae.
Group: Detarium.
Species Studied - Species in Genus: 1 studied; 1 in genus.

Description

Fruit: A legumelegume:
usually dry, dehiscent fruit derived from a single carpel that opens along two longitudinal sutures
; unilocular; 4.5–7 cm long; 3.5–4 cm wide; 1.1–1.4 cm thick; length less than twice as long as width; with deciduous androecial sheath; with deciduous corolla; with deciduous calyx; without orifice formed by curving of fruit or fruit segments; straight; not plicate; not twisted; asymmetrical; elliptic; not inflated; compressed; without beak; long tapered at apex to tapered at apex; aligned with longitudinal axis of fruit; rounded at base; aligned with longitudinal axis of fruit; with the apex and base uniform in texture; ligneous; seed chambers externally invisible; margin not constricted; margin without sulcus; margin plain; wing(s) absent; substipitate; with all layers dehiscing; splitting along suture(s). Dehiscence of valves along both sutures; passive. Replum invisible. Epicarp dull; monochrome; brown; with surface texture uniform; glabrous; eglandular; without spines; not smooth; with elevated features; not veined; not tuberculate; rugose; not exfoliating; without cracks; without embedded tissue, much thicker than epicarp, running from base to apex. Mesocarp present; surface not veined; 2-layered; without balsamic vesicles; with fibers; without reniform canals; with fibers embedded in mealy tissue over solid layer; ligneous. Endocarp present; visible; dull; opaque; monochrome; whitish brown; without adhering pieces of testa; nonseptate; not exfoliating; remaining fused to mesocarp and epicarp; entire. Seed(s) 1–2; length oblique to fruit length; neither overlapping nor touching; in 1 series. Funiculus 0.1–5 mm long; of 1 length only; thick; essentially straight. Aril present; fleshy; when fleshy type or shape not stated; entire; covering entire seed.

Seed: 28–30 mm long; 15–20 mm wide; 10–11 mm thick; not overgrown; not angular; symmetrical, or asymmetrical; oblong, or irregular; compressed; with surface smooth; without visible radicle and cotyledon lobes; without hilar sinus; without umbo on seed faces; without medial ridge on each face. Cuticle not exfoliating; not inflated; not wrinkled. Testa present; without pieces of adhering epicarp; not adhering to endocarp; free from endocarp; glossy; not modified by a bloom; colored; monochrome; dark brown to blackish brown; glabrous; not smooth; with elevated features; rugose; coriaceous (subcoriaceous). Pleurogram absent. Pseudopleurogram absent. Fracture lines absent. Rim absent. Wing(s) absent. Raphe not visible. Hilum present; fully concealed; concealed by funicular remnant; without faboid split; larger than punctiform; 0.1–2.5 mm long; with curved outline; elliptic; subapical to radicle tip; flush; not within corona, halo, or rim. Lens not discernible. Endosperm absent. Cotyledons smooth; both outer faces convex; both the same thickness; both more or less of equal length; not folded; margin entire 180 degrees from base of radicle; similar at apex; completely concealing radicle; split over radicle (and gaping); with the interface division terminating at base of radicle; without margins recessed; inner face flat; glabrous on inner face. Embryonic axis straight; oblique to length of seed; without a joint evident between the radicle and the cotyledons. Radicle differentiated from cotyledon; centered between cotyledons. Plumule rudimentary; glabrous.

Distribution

Borneo and Sarawak.

Old World; Indonesia and the Philippines (Borneo and Sarawak).

Generic Notes

"Debateably distinct from Copaifera" (4.43) (Cowan and Polhill, 1981a). Hou (1994, Blumea 38:313–330) synonmized Pseudosindora under Copaifera.

Tribal Notes

Tribe Detarieae

Bruneau et al. (2000) carried out extensive phylogenetic analyses of tribes Amherstieae and Detarieae. They concluded that they form a single monophyletic group. Therefore, they supported Polhill's (1995a, 1995b) decision to unite the two tribes.