Pachyelasma

Taxonomy

Pachyelasma H.A.T. Harms Bot. Jahrb. Syst. 49: 428. 28 Mai 1913.

Subfamily: Caesalpinioideae.
Phylogenetic Number: 1.1.39.
Tribe: Caesalpinieae.
Group: Dimorphandra.
Species Studied - Species in Genus: 1 studied; 1 in genus.

Description

Fruit: A legume; unilocular; 15–37 cm long; 3.5–5 cm wide; 1.5–2.5 cm thick; 2–9 times longer than wide; with deciduous androecial sheath; with deciduous corolla; with deciduous calyx; without orifice formed by curving of fruit or fruit segments; straight to curved; not plicate; not twisted; asymmetrical; oblong; not inflated; compressed; without beak; nearly tapered at apex; aligned with longitudinal axis of fruit; tapered at base to rounded at base; aligned with longitudinal axis of fruit; with the apex and base uniform in texture; ligneous; seed chambers externally invisible (faintly visible); margin not constricted; margin without sulcus; margin plain; wing(s) absent; nonstipitate; apparently indehiscent. Replum invisible. Epicarp dull; monochrome; blackish brown; with surface texture uniform; glabrous; eglandular; without spines; nearly smooth; not veined; not tuberculate; not exfoliating; without cracks; without embedded tissue, much thicker than epicarp, running from base to apex. Mesocarp present; surface not veined; 2-layered; without balsamic vesicles; without fibers; without reniform canals; with thick brown solid layer over solid layer to vitriol layer over solid layer (thin & tan); ligneous. Endocarp present; visible; dull; opaque; monochrome; tannish brown; without adhering pieces of testa; septate; not exfoliating; remaining fused to mesocarp and epicarp; entire. Seed(s) 10–18; length transverse to fruit length; neither overlapping nor touching; in 1 series. Funiculus 0.1–8 mm long; of 1 length only; filiform; straight and plicate. Aril absent.

Seed: 20–25 mm long; 8–13 mm wide; 5–6 mm thick; not overgrown; not angular; symmetrical; ovate to oblong; compressed; with surface smooth; without visible radicle and cotyledon lobes; without hilar sinus; without umbo on seed faces; without medial ridge on each face. Cuticle not exfoliating; not inflated; not wrinkled. Testa present; without pieces of adhering epicarp; not adhering to endocarp; free from endocarp; dull; not modified by a bloom; colored; monochrome; brown; glabrous; smooth; osseous. Pleurogram present; 90 %. Pseudopleurogram absent. Fracture lines present; reticulate. Rim absent. Wing(s) absent. Raphe not visible. Hilum present; fully concealed; concealed by funicular remnant; without faboid split; punctiform; subapical to radicle tip; flush; not within corona, halo, or rim. Lens discernible; 0.1–0.5 mm long; with margins curved; elliptic; not in groove of raphe; flush (or nearly so); dissimilar color from testa; black (sometimes center tan); not within corona, halo, or rim. Endosperm present; thick; not pluglike and not resembling tip of radicle; adnate to embryo. Cotyledons smooth; both outer faces convex; both the same thickness; both more or less of equal length; not folded; margin entire 180 degrees from base of radicle; similar at apex; partially concealing radicle; with lobes; with lobes not touching; with the interface division terminating at base of radicle; without margins recessed; inner face flat; glabrous on inner face. Embryonic axis straight; parallel to length of seed; without a joint evident between the radicle and the cotyledons. Radicle differentiated from cotyledon; centered between cotyledons. Plumule moderately developed; glabrous.

Distribution

South Nigeria to Central African Republic, Zaire, and Gabon.

Old World; Africa (South Nigeria to Central African Republic, Zaire, and Gabon).
 Fruit and seed:  P. tessmannii  (H.A.T. Harms) H.A.T. Harms - top left seeds in situ, top right transverse fruit section [after Harms, 1913], bottom fruit with seed in situ.
Fruit and seed: P. tessmannii (H.A.T. Harms) H.A.T. Harms - top left seeds in situ, top right transverse fruit section [after Harms, 1913], bottom fruit with seed in situ.
 Seed, cotyledon, embryo, and testa:  P. tessmannii  (H.A.T. Harms) H.A.T. Harms - top left cotyledon investing exposed radicle (L) and embryonic axis (R), bottom far left seed topography, bottom left center seeds, testa SEMs.
Seed, cotyledon, embryo, and testa: P. tessmannii (H.A.T. Harms) H.A.T. Harms - top left cotyledon investing exposed radicle (L) and embryonic axis (R), bottom far left seed topography, bottom left center seeds, testa SEMs.