Moldenhawera

Taxonomy

Moldenhawera H.A. Schrader Gött. Gel. Anz. 1: 718. 1802.

Subfamily: Caesalpinioideae.
Phylogenetic Number: 1.1.18.
Tribe: Caesalpinieae.
Group: Peltophorum.
Species Studied - Species in Genus: 3 studied; 6 in genus.

Description

Fruit: A legume; unilocular; 7–20 cm long; 1.8–2.5 cm wide; 0.2–4 cm thick; 2–9 times longer than wide; with deciduous androecial sheath; with deciduous corolla; with deciduous calyx; without orifice formed by curving of fruit or fruit segments; straight to curved; not plicate; not twisted; symmetrical; oblong; not inflated; flattened; without beak; long tapered at apex to tapered at apex; aligned with longitudinal axis of fruit; short tapered at base to rounded at base; aligned with longitudinal axis of fruit; with the apex and base uniform in texture; coriaceous; seed chambers externally invisible, or visible; margin constricted, or not constricted; margin without sulcus; margin plain; wing(s) absent; substipitate to nonstipitate; with all layers dehiscing; splitting along suture(s). Dehiscence of valves along 1 suture; passive. Replum invisible. Epicarp dull; monochrome; brown, or black (reddish); with surface texture uniform; golden to reddish pubescent and indurate; with hairs erect; with simple hairs; eglandular; without spines; not smooth; not veined; not tuberculate; not exfoliating; with cracks; cracking longitudinal to fruit length; without embedded tissue, much thicker than epicarp, running from base to apex. Mesocarp present; surface not veined; 1-layered; without balsamic vesicles; without fibers; without reniform canals; solid; ligneous. Endocarp present; visible; dull; opaque; monochrome; brown; without adhering pieces of testa; nonseptate; not exfoliating; remaining fused to mesocarp and epicarp; entire. Seed(s) 3–11; length transverse to fruit length; neither overlapping nor touching; in 1 series. Funiculus 0.1–0.7 mm long; of 1 length only; flattened; straight. Aril absent.

Seed: 12–15 mm long; 9–11 mm wide; 1 mm thick; not overgrown; not angular; symmetrical; ovate; flattened; with surface smooth; without visible radicle and cotyledon lobes; without hilar sinus; without umbo on seed faces; without medial ridge on each face. Cuticle not exfoliating; not inflated; not wrinkled. Testa present; without pieces of adhering epicarp; not adhering to endocarp; free from endocarp; glossy; not modified by a bloom; colored; monochrome; brown; glabrous; smooth; osseous. Pleurogram absent. Pseudopleurogram absent. Fracture lines absent. Rim absent. Wing(s) absent. Raphe not visible. Hilum present; visible; without faboid split; larger than punctiform; 0.1–1.5 mm long; with curved outline, or straight outline; elliptic; apical at apex of radicle tip; flush; not within corona, halo, or rim. Lens not discernible. Endosperm present; thin; not pluglike and not resembling tip of radicle; adnate to testa. Cotyledons smooth; both outer faces convex; both the same thickness; both more or less of equal length; not folded; margin entire 180 degrees from base of radicle; similar at apex; partially concealing radicle; with lobes (1 side auriculate because of slight radicle inflection); with lobes not touching; with the interface division terminating at base of radicle; without margins recessed; inner face flat; glabrous on inner face. Embryonic axis straight; slightly oblique to length of seed; without a joint evident between the radicle and the cotyledons. Radicle differentiated from cotyledon; centered between cotyledons. Plumule rudimentary; glabrous.

Distribution

Brazil and Venezuela.

New World; South America (Brazil and Venezuela); Brazil.

Generic Notes

The genus is usually spelled Moldenhauera, but Polhill and Vidal (1981) observed "perhaps more correctly spelt Moldenhawera." Yakovlev (1975) spelled the genus Moldenhawera in his monograph.

 Fruit and seed:  M. cuprea  Pohl - bottom seeds in situ;  M. floribunda  H.A. Schrader - top fruit.
Fruit and seed: M. cuprea Pohl - bottom seeds in situ; M. floribunda H.A. Schrader - top fruit.
 Seed, cotyledon, embryo, and testa:  M. cuprea  Pohl - bottom far left seed topography, top left cotyledon cordate and investing nearly exposed radicle (L) and embryonic axis (R), testa SEMs;  M. floribunda  H.A. Schrader - bottom left center seed.
Seed, cotyledon, embryo, and testa: M. cuprea Pohl - bottom far left seed topography, top left cotyledon cordate and investing nearly exposed radicle (L) and embryonic axis (R), testa SEMs; M. floribunda H.A. Schrader - bottom left center seed.