Hammatolobium

Taxonomy

Hammatolobium E. Fenzl Pugillus Pl. Nov. 1 ('Hamatolobium'). t. 1. Mai-Jun 1842(?).

Subfamily: Faboideae.
Phylogenetic Number: 3.13.17.
Tribe: Loteae.
Species Studied - Species in Genus: 2 studied; 2 in genus.

Description

Fruit: A loment (or a loment segment); 2–6 cm long; 0.15–0.4 cm wide; 0.2 cm thick; more than 9 times longer than wide; with deciduous androecial sheath; with deciduous corolla; with persistent calyx; with calyx shorter than fruit; without orifice formed by curving of fruit or fruit segments; straight; not plicate; not twisted; symmetrical; linear and moniliform; not inflated; compressed; with beak; straight; with solid beak the same color and texture as fruit; short tapered at apex; oblique with longitudinal axis of fruit; short tapered at base; aligned with longitudinal axis of fruit; with the apex and base uniform in texture; coriaceous; seed chambers externally visible; seed chambers with the raised seed chambers not torulose; margin constricted; margin constricted along both margins; margin without sulcus; margin plain; wing(s) absent; substipitate; indehiscent. Replum invisible. Loment indehiscent; segments (articles) inconspicuous; segments (articles) 3–6 mm long; segments (articles) widest across seed area; segments (articles) with apical 1 different shape than middle one(s), or basal 1 different shape than middle one(s); segments (articles) oblong. Epicarp dull; monochrome; dark brown; with surface texture uniform; pubescent and indurate; with hairs erect; with 1 type of pubescence; pilose, or villous; with pubescence golden; with pubescence uniformly distributed; with simple hairs; pliable; with hair bases plain; eglandular; without spines; smooth, or not smooth; with elevated features; veined; reticulately veined; not tuberculate; not exfoliating; without cracks; without embedded tissue, much thicker than epicarp, running from base to apex. Mesocarp present; thin; surface not veined; 1-layered; without balsamic vesicles; without fibers; without reniform canals; solid; coriaceous. Endocarp present; visible; glossy; opaque; monochrome; white; smooth; without adhering pieces of testa; septate; with septa thicker than paper, firm; with septa eglandular; coriaceous; not exfoliating; remaining fused to mesocarp and epicarp; without wings; entire. Seed(s) 5–11 (estimated); length parallel with fruit length; neither overlapping nor touching; in 1 series. Funiculus of 1 length only; thick; straight. Aril absent.

Seed: 2.5 mm long; 2.3 mm wide; 1 mm thick; not overgrown; not angular; asymmetrical; nearly circular; compressed; without visible radicle and cotyledon lobes; without hilar sinus; without umbo on seed faces; without medial ridge on each face. Cuticle not exfoliating; not inflated; not wrinkled. Testa present; without pieces of adhering epicarp; not adhering to endocarp; free from endocarp; dull; not modified by a bloom; colored; monochrome; reddish brown; glabrous; smooth; coriaceous. Pleurogram absent. Pseudopleurogram absent. Fracture lines absent. Rim absent. Wing(s) absent. Raphe not visible. Hilum present; fully concealed; concealed by funicular remnant; without faboid split; punctiform; between cotyledon and radicle lobe; recessed; not within corona, halo, or rim. Lens discernible; with margins curved; elliptic; not in groove of raphe; adjacent to hilum; mounded; dissimilar color from testa; darker than testa; black; not within corona, halo, or rim. Endosperm present; thin; not pluglike and not resembling tip of radicle; covering entire embryo; adnate to testa. Cotyledons smooth; both outer faces convex; both the same thickness; both more or less of equal length; not folded; margin entire 180 degrees from base of radicle; similar at apex; not concealing radicle; entire over radicle; without lobes; with the interface division terminating at base of radicle; without margins recessed; yellow; inner face flat; glabrous on inner face. Embryonic axis deflexed; oblique to length of seed; without a joint evident between the radicle and the cotyledons. Radicle differentiated from cotyledon; linear; oblique to cotyledons; centered between cotyledons; 1/2 to nearly length of cotyledons. Plumule rudimentary; glabrous.

Distribution

One species at either end of the Mediterranean region.

Old World; Europe (Mediterranean).

Generic Notes

Polhill (1981l) noted that this genus seems to be more closely related to Anthyllis (13.02) and Lotus (13.07) in the Loteae than to Coronilla (13.11).

Tribal Notes

Tribe Loteae

In 1981, Polhill (1981k) accepted much broader generic circumscriptions in tribe Loteae, and only accepted four genera in the tribe: Cytisopsis, Anthyllis (13.02), Hymenocarpus (13.04), and Lotus (13.07). In his most recent classification of Fabaceae (Polhill, 1994a, 1994b), he combined tribes Loteae and Coronilleae and accepted six segregate genera in Loteae, s.s.: Tripodion (13.03), Dorycnopsis (13.05), Dorycnium (13.06), Podolotus J.F. Royle (13.08), Pseudolotus K.H. Rechinger (13.09), and Vermifrux (13.10).