Errazurizia

Taxonomy

Errazurizia R.A. Philippi Anales Univ. Chile 1872: 688. Nov 1872.

Subfamily: Faboideae.
Phylogenetic Number: 3.6.05.
Tribe: Amorpheae.
Species Studied - Species in Genus: 4 studied; 4 in genus.

Description

Fruit: A legume; unilocular; 0.4–1.1 cm long; 0.45–0.8 cm wide; 0.5–0.6 cm thick; length less than twice as long as width to 2–9 times longer than wide; with deciduous androecial sheath; with deciduous corolla; with persistent calyx; with calyx shorter than fruit; without orifice formed by curving of fruit or fruit segments; straight; not plicate; not twisted; asymmetrical, or symmetrical; elliptic, or ovate; with 1 straight and 1 curved suture; widest near middle or D-shaped; not inflated; compressed; without beak, or with beak; straight; with solid beak the same color and texture as fruit; rounded at apex; oblique with longitudinal axis of fruit; tapered at base; aligned with longitudinal axis of fruit; with the apex and base differing in texture; chartaceous; seed chambers externally visible; margin not constricted; margin without sulcus; margin plain; wing(s) absent; nonstipitate; indehiscent. Replum invisible. Epicarp dull; monochrome (except for glandular dots); brown, or gray (because of gray hairs); with surface texture uniform; pubescent and indurate; with hairs erect; with 1 type of pubescence; villous; with pubescence gray; with pubescence uniformly distributed; with simple hairs; pliable; with hair bases plain; glandular; with glandular dots; limited to a portion of fruit; upper 1/2 glandular and lower 1/2 eglandular; without spines; not smooth; with elevated features; not veined; not tuberculate; wrinkled; not exfoliating; without cracks; without embedded tissue, much thicker than epicarp, running from base to apex. Mesocarp absent. Endocarp present; visible; dull; opaque; monochrome; brown; smooth; without adhering pieces of testa; nonseptate; chartaceous; not exfoliating; remaining fused to epicarp; without wings; entire. Seed(s) 1; length parallel with fruit length. Funiculus filiform; straight. Aril absent.

Seed: 3–7.5 mm long; 2.7–5 mm wide; 2–4.5 mm thick; not overgrown; not angular; symmetrical (except for hilum); elliptic; compressed; without visible radicle and cotyledon lobes; without hilar sinus; without umbo on seed faces; without medial ridge on each face. Cuticle not exfoliating; not inflated; not wrinkled. Testa present; without pieces of adhering epicarp; not adhering to endocarp; free from endocarp; dull (with splotches of glandular residue which are shiny); not modified by a bloom; colored; monochrome; black, or brown, or tan, or purple; glabrous; smooth; coriaceous. Pleurogram absent. Pseudopleurogram absent. Fracture lines absent. Rim absent. Wing(s) absent. Raphe visible; from lens to base of seed and terminating; not bifurcating; darker than testa; black; deeply recessed. Hilum present; fully concealed; concealed by funicular remnant; without faboid split; punctiform; marginal according to radicle tip; deeply recessed; not within corona, halo, or rim. Lens not discernible. Endosperm present; thick; not pluglike and not resembling tip of radicle; covering entire embryo; adnate to testa. Cotyledons smooth; both outer faces convex; both the same thickness; both more or less of equal length; not folded; margin entire 180 degrees from base of radicle; similar at apex; partially concealing radicle; split over radicle; with lobes; with lobes touching (auriculate); with basal groin formed by lobes; with the interface division terminating at base of radicle; without margins recessed; green, or tan; inner face flat; glabrous on inner face. Embryonic axis deflexed; oblique to length of seed; without a joint evident between the radicle and the cotyledons. Radicle differentiated from cotyledon; bulbose, or linear; lobe tip curved; deflexed and parallel to cotyledon width; not centered between cotyledons (radicle outside 1 cotyledon and inside other, therefore junctions for each cotyledon different); less than 1/2 length of cotyledons. Plumule moderately developed; glabrous.

Distribution

Three species in southwest United States and northwestern Mexico, And one species along the coast of Chile.

New World; southwestern United States, Mexico, and South America (coastal Chile, Mexico (northwestern)).

Generic Notes

Barneby (1977) monographed Errazurizia.

Tribal Notes

Tribe Amorpheae

Traditionally this tribe has been called Amorpheae. Reveal (1997) reported that the name Daleeae was published before the name Amorpheae. Following the International Code of Botanical Nomenclature (Greuter et al., 1994), the oldest name for a taxon must be used, so Reveal suggested that this tribe should called Daleeae. In 1999 Reveal (1999) reversed himself, so that the tribe remains the Amorpheae.

 Fruit and seed:  E.  spp. - fruits with calyx and seeds.
Fruit and seed: E. spp. - fruits with calyx and seeds.
 Cotyledon, embryo, and testa:  E. rotundata  (E.O. Wooton) R.C. Barneby - embryo, cotyledons, and testa SEMs.
Cotyledon, embryo, and testa: E. rotundata (E.O. Wooton) R.C. Barneby - embryo, cotyledons, and testa SEMs.