Endertia

Taxonomy

Endertia C.G.G.J. van Steenis & H.C.D. De Wit Bull. Jard. Bot. Buitenzorg ser. 3. 17: 323. Dec 1947.

Subfamily: Caesalpinioideae.
Phylogenetic Number: 1.4.18.
Tribe: Detarieae.
Group: Hymenostegia.
Species Studied - Species in Genus: 1 studied; 1 in genus.

Description

Fruit: A legume; unilocular; 26–27 cm long; 4–6 cm wide; 0.8–1 cm thick; 2–9 times longer than wide; with deciduous androecial sheath; with deciduous corolla; with deciduous calyx; without orifice formed by curving of fruit or fruit segments; straight; not plicate; not twisted; symmetrical, or asymmetrical; with 1 straight and 1 curved suture; widest near apex ("to scimitar-shaped" with edited comment "no value"); not inflated; flattened; without beak; dorsal suture short long tapered at apex; oblique with longitudinal axis of fruit (slightly); tapered at base; oblique with longitudinal axis of fruit; with the apex and base uniform in texture; ligneous; seed chambers externally invisible; margin not constricted; margin without sulcus; margin plain; wing(s) absent; not stated; tardily with all layers dehiscing; splitting along suture(s). Dehiscence of valves along both sutures; active; with valves separately enrolling. Replum invisible. Epicarp dull; monochrome; brown; with surface texture uniform; glabrous; eglandular; without spines; not smooth; with elevated features; when young veined; reticulately veined; not tuberculate; lenticular (becoming smooth with numerous narrowly elliptic lenticels); not exfoliating; without cracks; without embedded tissue, much thicker than epicarp, running from base to apex. Mesocarp present; surface not veined; 2-layered; without balsamic vesicles; with fibers; without reniform canals; with fibers spongy tissue with transverse over solid layer; ligneous. Endocarp present; visible; dull; opaque; monochrome; brown; without adhering pieces of testa; nonseptate; not exfoliating; remaining fused to mesocarp and epicarp; entire. Seed(s) 3–5; length oblique to fruit length; neither overlapping nor touching; in 1 series. Funiculus 0.1–3 mm long; of 1 length only; thick; straight. Aril absent.

Seed: 31–42 mm long; 21–25 mm wide; 8–10 mm thick; not overgrown; not angular; symmetrical; circular to oblong; compressed; with surface smooth; without visible radicle and cotyledon lobes; without hilar sinus; without umbo on seed faces; without medial ridge on each face. Cuticle not exfoliating; not inflated; not wrinkled. Testa present; without pieces of adhering epicarp; not adhering to endocarp; free from endocarp; glossy; not modified by a bloom; colored; monochrome; reddish to blackish brown; glabrous; smooth, or not smooth; with elevated features; rugose; not stated. Pleurogram absent. Pseudopleurogram absent. Fracture lines absent. Rim absent. Wing(s) absent. Raphe not visible. Hilum present; fully concealed; concealed by funicular remnant; without faboid split; larger than punctiform; not stated; with curved outline; circular; marginal according to radicle tip (?, stated as nearly lateral); recessed; not within corona, halo, or rim. Lens not discernible. Cotyledons smooth; both outer faces convex; both the same thickness; both more or less of equal length; not folded; margin entire 180 degrees from base of radicle; similar at apex; completely concealing radicle; notched at radicle and split over radicle; with the interface division terminating at base of radicle; without margins recessed; inner face flat; glabrous on inner face. Embryonic axis straight; oblique to length of seed; without a joint evident between the radicle and the cotyledons. Radicle differentiated from cotyledon; centered between cotyledons. Plumule rudimentary; glabrous.

Distribution

Borneo.

Old World; Indonesia and the Philippines (Borneo).

Generic Notes

De Wit (1947a) studied Endertia.

Tribal Notes

Tribe Detarieae

Bruneau et al. (2000) carried out extensive phylogenetic analyses of tribes Amherstieae and Detarieae. They concluded that they form a single monophyletic group. Therefore, they supported Polhill's (1995a, 1995b) decision to unite the two tribes.
 Fruit and seed:  E. spectabilis  C.G.G.J. van Steenis & H.C.D. de Wit - top fruit, bottom seed in situ.
Fruit and seed: E. spectabilis C.G.G.J. van Steenis & H.C.D. de Wit - top fruit, bottom seed in situ.
 Seed, cotyledon, embryo, and testa:  E. spectabilis  C.G.G.J. van Steenis & H.C.D. de Wit - top left cotyledon notched and split (L) and embryonic axis (R), bottom far left seed topography, bottom left center seed, testa SEMs.
Seed, cotyledon, embryo, and testa: E. spectabilis C.G.G.J. van Steenis & H.C.D. de Wit - top left cotyledon notched and split (L) and embryonic axis (R), bottom far left seed topography, bottom left center seed, testa SEMs.