Balsamocarpon

Taxonomy

Balsamocarpon D. Clos In C. Gay, Hist. Chile, Bot. 2: 226. t. 20. Ante Mai 1847.

Subfamily: Caesalpinioideae.
Phylogenetic Number: 1.1.33.
Tribe: Caesalpinieae.
Group: Caesalpinia.
Species Studied - Species in Genus: 1 studied; 1 in genus.

Description

Fruit: A legume; unilocular; 3.8–4.1 cm long; 1.2–1.5 cm wide; 0.6–1 cm thick; 2–9 times longer than wide; with deciduous androecial sheath; with deciduous corolla; with deciduous calyx; without orifice formed by curving of fruit or fruit segments; straight; not plicate; not twisted; asymmetrical; oblong; not inflated; compressed to subtriangular; without beak; tapered at apex; aligned with longitudinal axis of fruit; short tapered at base; aligned with longitudinal axis of fruit; with the apex and base uniform in texture; ligneous; seed chambers externally visible; margin not constricted; margin without sulcus; margin plain; wing(s) absent; nonstipitate; indehiscent. Replum invisible. Epicarp dull; monochrome; brown; with surface texture uniform; pubescent and indurate; with hairs erect; gray eglandular, or glandular; with glandular red hairs; without spines; not smooth; with elevated features; not veined; not tuberculate; rugose; not exfoliating; without cracks; without embedded tissue, much thicker than epicarp, running from base to apex. Mesocarp present; surface not veined; 2-layered; without balsamic vesicles; with fibers; without reniform canals; with fibers transverse reticulate over vitriol layer; ligneous. Endocarp present; visible; dull; opaque; monochrome; white; without adhering pieces of testa; subseptate; not exfoliating; remaining fused to mesocarp and epicarp; entire. Seed(s) 4–5; length transverse to fruit length; neither overlapping nor touching; in 1 series. Funiculus 0.1–1.5 mm long; of 1 length only; filiform; S-curved. Aril absent.

Seed: 7.5–8.5 mm long; 7.5–8 mm wide; 4.5–6 mm thick; not overgrown; not angular; symmetrical; circular; compressed; with surface smooth; without visible radicle and cotyledon lobes; without hilar sinus; without umbo on seed faces; without medial ridge on each face. Cuticle not exfoliating; not inflated; not wrinkled. Testa present; without pieces of adhering epicarp; not adhering to endocarp; free from endocarp; dull; not modified by a bloom; colored; monochrome; brown; glabrous; not smooth; with elevated features; shagreen; osseous. Pleurogram absent. Pseudopleurogram absent. Fracture lines absent. Rim absent. Wing(s) absent. Raphe not visible. Hilum present; fully concealed; concealed by funicular remnant; without faboid split; punctiform; apical at apex of radicle tip; recessed (depression slitlike); not within corona, halo, or rim. Lens discernible; 0.3 mm long; with margins curved; circular; not in groove of raphe; recessed; dissimilar color from testa; black; not within corona, halo, or rim. Endosperm present; thick; not pluglike and not resembling tip of radicle; adnate to testa. Cotyledons smooth; both outer faces convex; both the same thickness; both more or less of equal length; not folded; margin entire 180 degrees from base of radicle; similar at apex; partially concealing radicle; with lobes; with lobes overlapping; with basal groin formed by lobes; with the interface division terminating at base of radicle; without margins recessed; inner face flat; glabrous on inner face. Embryonic axis straight; parallel to length of seed; without a joint evident between the radicle and the cotyledons. Radicle differentiated from cotyledon; centered between cotyledons. Plumule rudimentary; glabrous.

Distribution

Chile.

New World; South America (Chile).
 

Generic Notes

Polhill and Vidal (1981) regarded this genus as "a very doubtful segregate of Caesalpinia, distinguished solely by the pinnate leaves, a feature rare in Caesalpinia as traditionally circumscribed (there are several pinnate leaved Caesalpinias in Cuba)".
 Fruit and seed:  B. brevifolium  D. Clos - top eroded fruit, left seed in situ, right mesocarp.
Fruit and seed: B. brevifolium D. Clos - top eroded fruit, left seed in situ, right mesocarp.
 Seed, cotyledon, embryo, and testa:  B. brevifolium  D. Clos - top left cotyledon split and basal groined and concealing all but radicle tip (L) and embryonic axis (R), bottom far left seed topography, bottom left center seeds, testa SEMs.
Seed, cotyledon, embryo, and testa: B. brevifolium D. Clos - top left cotyledon split and basal groined and concealing all but radicle tip (L) and embryonic axis (R), bottom far left seed topography, bottom left center seeds, testa SEMs.