Archidendropsis

Taxonomy

Archidendropsis I.C. Nielsen Fl. Nouv.-Calédonie 12: 66. 1983 (post 31 Mai).

Subfamily: Mimosoideae.
Phylogenetic Number: 2.5.17.
Tribe: Ingeae.
Species Studied - Species in Genus: 3 studied; 14 in genus.

Description

Fruit: A legume; unilocular; 4.2–18 cm long; 0.5–3 cm wide; 0.5–2.5 cm thick; 2–9 times longer than wide, or more than 9 times longer than wide; with deciduous androecial sheath; with deciduous corolla; with deciduous calyx; without orifice formed by curving of fruit or fruit segments; straight to curved; not plicate; not twisted; asymmetrical to symmetrical; linear to oblong, or C-shaped; with both sutures parallelly curved; not inflated; flattened to compressed; without beak; short tapered at apex to rounded at apex; aligned with longitudinal axis of fruit; short tapered at base; aligned with longitudinal axis of fruit; with the apex and base uniform in texture; chartaceous to ligneous; seed chambers externally visible to invisible; margin constricted; margin constricted along both margins to slightly constricted along both margins; margin without sulcus; margin plain; wing(s) absent; substipitate; with all layers dehiscing; splitting along suture(s). Dehiscence of valves along both sutures (and scissoring apart, remaining attached to sutures); apical and down; passive. Replum invisible. Epicarp dull to glossy; monochrome; yellowish brown to dark brown; with surface texture uniform; pubescent and indurate to glabrate, or glabrous; with hairs erect; with simple hairs; eglandular; without spines; not smooth; with elevated features; veined; reticulately veined; not tuberculate; not exfoliating; without cracks; without embedded tissue, much thicker than epicarp, running from base to apex. Mesocarp absent. Endocarp present; visible; dull; opaque; monochrome; tan to brown; smooth; without adhering pieces of testa; nonseptate; not exfoliating; entire. Seed(s) 11–13; length parallel with fruit length; neither overlapping nor touching; in 1 series. Funiculus 4–5 mm long; of 1 length only; filiform; curved to plicate. Aril absent.

Seed: 5–28 mm long; 5.5–17 mm wide; 0.5–1 mm thick; not overgrown; not angular; symmetrical; oblong; flattened; with surface smooth; without visible radicle and cotyledon lobes; without hilar sinus; without umbo on seed faces; without medial ridge on each face. Cuticle not exfoliating; not inflated; not wrinkled. Testa present; without pieces of adhering epicarp; not adhering to endocarp; free from endocarp; glossy; not modified by a bloom; colored; monochrome; dark brown; glabrous; not smooth; with elevated features; rugose; chartaceous. Pleurogram absent. Pseudopleurogram absent. Fracture lines absent. Rim absent. Wing(s) present; at both ends (up to 2 mm wide). Raphe not visible. Hilum present; fully concealed; concealed by funicular remnant; without faboid split; punctiform; apical according to radicle tip but marginal according to seed length; flush (but in notch); not within corona, halo, or rim. Lens not discernible. Endosperm absent. Cotyledons smooth; both outer faces flat; both the same thickness; both more or less of equal length; not folded; margin entire 180 degrees from base of radicle; similar at apex; not concealing radicle; notched at radicle; with the interface division terminating at base of radicle; without margins recessed; inner face flat; glabrous on inner face. Embryonic axis straight; perpendicular to length of seed; without a joint evident between the radicle and the cotyledons. Radicle differentiated from cotyledon; straight with embryonic axis; centered between cotyledons. Plumule well developed; glabrous.

Distribution

New Guinea, Soloman Islands to New Caledonia, and northeastern Australia.

Old World; Indonesia and the Philippines, Pacific, and Australia (northeastern).
 

Generic Notes

The seed of A. streptocarpa is similar in structure to the seed of Anadenathera colubrina (J.M. da C. Vellozo) J.P.M. Brenan. Nielsen et al. (1983) monographed the genus and is the source of the number of species in the genus, not Nielsen (1981a). Fruits of A. basaltica (F.J.H. von Mueller) I.C. Nielsen are glandular (Nielsen et al., 1983). Nielsen et al. (1983a, 1983b) revised the genus, and their species count and distribution are used.