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A dead bee cluster likely due to starvation; photo by Jenie F. Warner A dead bee cluster likely due to starvation; photo by Jenie F. Warner
Deadouts
A dead bee cluster likely due to starvation; photo by Jenie F. Warner
Winter deadout with evidence of dysentery; photo by Wayne Dailey Winter deadout with evidence of dysentery; photo by Wayne Dailey
Deadouts
Winter deadout with evidence of dysentery; photo by Wayne Dailey
Dead winter cluster with mold; photo by The BeeMD photo collection Dead winter cluster with mold; photo by The BeeMD photo collection
Deadouts
Dead winter cluster with mold; photo by The BeeMD photo collection
Winter deadout of small cluster; photo by Dewey M. Caron Winter deadout of small cluster; photo by Dewey M. Caron
Deadouts
Winter deadout of small cluster; photo by Dewey M. Caron

Cannibalized or non-emerged brood in deadout; photo by Dewey M. Caron Cannibalized or non-emerged brood in deadout; photo by Dewey M. Caron
Deadouts
Cannibalized or non-emerged brood in deadout; photo by Dewey M. Caron
Apiary with dearth of floral sources; photo by Dewey M. Caron Apiary with dearth of floral sources; photo by Dewey M. Caron
Dearth
Apiary with dearth of floral sources; photo by Dewey M. Caron
Apiary with dearth of nectar and pollen resources; photo by Dewey M. Caron Apiary with dearth of nectar and pollen resources; photo by Dewey M. Caron
Dearth
Apiary with dearth of nectar and pollen resources; photo by Dewey M. Caron
Large commercial apiary under dearth conditions; photo by Dewey M. Caron Large commercial apiary under dearth conditions; photo by Dewey M. Caron
Dearth
Large commercial apiary under dearth conditions; photo by Dewey M. Caron

Defensive Africanized honey bees; photo by Maryann Frazier Defensive Africanized honey bees; photo by Maryann Frazier
Defensive colony
Defensive Africanized honey bees; photo by Maryann Frazier
Defensive (Africanized) bees about bee veil; photo by Maryann Frazier Defensive (Africanized) bees about bee veil; photo by Maryann Frazier
Defensive colony
Defensive (Africanized) bees about bee veil; photo by Maryann Frazier
Defensive bees flying directly to bee veil; they're attracted to carbon dioxide; photo by Dewey M. Caron Defensive bees flying directly to bee veil; they're attracted to carbon dioxide; photo by Dewey M. Caron
Defensive colony
Defensive bees flying directly to bee veil; they're attracted to carbon dioxide; photo by Dewey M. Caron
Adult with DWV (right), adult with normal wings (left); photo by The BeeMD photo collection Adult with DWV (right), adult with normal wings (left); photo by The BeeMD photo collection
Deformed wing virus
Adult with DWV (right), adult with normal wings (left); photo by The BeeMD photo collection

Bee in the center has DWV; photo by The BeeMD photo collection Bee in the center has DWV; photo by The BeeMD photo collection
Deformed wing virus
Bee in the center has DWV; photo by The BeeMD photo collection
Worker adult with deformed wings and two Varroa mites on its thorax; photo by The BeeMD photo collection Worker adult with deformed wings and two Varroa mites on its thorax; photo by The BeeMD photo collection
Deformed wing virus
Worker adult with deformed wings and two Varroa mites on its thorax; photo by The BeeMD photo collection
Adult with DWV; photo by The BeeMD photo collection Adult with DWV; photo by The BeeMD photo collection
Deformed wing virus
Adult with DWV; photo by The BeeMD photo collection
Adult worker with DWV and perhaps also CWV; photo by Robert Snyder Adult worker with DWV and perhaps also CWV; photo by Robert Snyder
Deformed wing virus
Adult worker with DWV and perhaps also CWV; photo by Robert Snyder

Workers with deformed wings; photo by Penn State University Workers with deformed wings; photo by Penn State University
Deformed wing virus
Workers with deformed wings; photo by Penn State University
Bee in the center (arrow) with DWV. Note queen in the upper right; photo by Robert Snyder Bee in the center (arrow) with DWV. Note queen in the upper right; photo by Robert Snyder
Deformed wing virus
Bee in the center (arrow) with DWV. Note queen in the upper right; photo by Robert Snyder
Bee with DWV; photo by Paul Andersen Bee with DWV; photo by Paul Andersen
Deformed wing virus
Bee with DWV; photo by Paul Andersen
Shaking behavior as a signal; illustration courtesy of Tom Seeley Shaking behavior as a signal; illustration courtesy of Tom Seeley
Dorso-ventral shaking behavior
Shaking behavior as a signal; illustration courtesy of Tom Seeley

Colonies in apiary distinctly painted to reduce drifting; photo Dewey M. Caron Colonies in apiary distinctly painted to reduce drifting; photo Dewey M. Caron
Drifting
Colonies in apiary distinctly painted to reduce drifting; photo Dewey M. Caron
Drifting of bees occurred to the two end colonies; photo by Dewey M. Caron Drifting of bees occurred to the two end colonies; photo by Dewey M. Caron
Drifting
Drifting of bees occurred to the two end colonies; photo by Dewey M. Caron
Drifting of bees to the colony at the end; illustration courtesy Dave Cushman's website Drifting of bees to the colony at the end; illustration courtesy Dave Cushman's website
Drifting
Drifting of bees to the colony at the end; illustration courtesy Dave Cushman's website
Dysentery on frame top bars; photo by The BeeMD photo collection Dysentery on frame top bars; photo by The BeeMD photo collection
Dysentery
Dysentery on frame top bars; photo by The BeeMD photo collection

Dysentery on the front of colony; photo by Dewey M. Caron Dysentery on the front of colony; photo by Dewey M. Caron
Dysentery
Dysentery on the front of colony; photo by Dewey M. Caron
Dysentery on top bars; photo by Wayne Dailey Dysentery on top bars; photo by Wayne Dailey
Dysentery
Dysentery on top bars; photo by Wayne Dailey
Emergency queen cells; photo by The BeeMD photo collection Emergency queen cells; photo by The BeeMD photo collection
Emergency queen cells
Emergency queen cells; photo by The BeeMD photo collection
Emergency queen cells; photo by Penn State University Emergency queen cells; photo by Penn State University
Emergency queen cells
Emergency queen cells; photo by Penn State University

Emergency queen cells; photo by Dewey M. Caron Emergency queen cells; photo by Dewey M. Caron
Emergency queen cells
Emergency queen cells; photo by Dewey M. Caron
Entombed cells of contaminated pollen; photo by BeeInformed Partnership Entombed cells of contaminated pollen; photo by BeeInformed Partnership
Entombed pollen
Entombed cells of contaminated pollen; photo by BeeInformed Partnership
Decomposing larvae in several brood cells and twisted larvae in cells in upper right characteristic of EFB; photo by Robert Snyder Decomposing larvae in several brood cells and twisted larvae in cells in upper right characteristic of EFB; photo by Robert Snyder
European foulbrood
Decomposing larvae in several brood cells and twisted larvae in cells in upper right characteristic of EFB; photo by Robert Snyder
Larvae infected with EFB; photo by Robert Snyder Larvae infected with EFB; photo by Robert Snyder
European foulbrood
Larvae infected with EFB; photo by Robert Snyder