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Capped queen cells; photo by Elaine Timm Capped queen cells; photo by Elaine Timm
Queen cup
Capped queen cells; photo by Elaine Timm
Numerous queen cups; photo by Charles Vanden Heuvel Numerous queen cups; photo by Charles Vanden Heuvel
Queen cup
Numerous queen cups; photo by Charles Vanden Heuvel
Queen cups hanging vertically from bottom bars of top box; photo by Dewey M. Caron Queen cups hanging vertically from bottom bars of top box; photo by Dewey M. Caron
Queen cup
Queen cups hanging vertically from bottom bars of top box; photo by Dewey M. Caron
Queen lacking retinue could signify queen failure; photo by Kathy Keatley Garvey Queen lacking retinue could signify queen failure; photo by Kathy Keatley Garvey
Queen failure
Queen lacking retinue could signify queen failure; photo by Kathy Keatley Garvey

Newly emerged virgin queen; photo by Dan Borkoski Newly emerged virgin queen; photo by Dan Borkoski
Queen failure
Newly emerged virgin queen; photo by Dan Borkoski
Queen that fainted; photo by Zachary Huang, beetography.com Queen that fainted; photo by Zachary Huang, beetography.com
Queen fainting
Queen that fainted; photo by Zachary Huang, beetography.com
Queen on frame of ripening nectar above queen excluder; photo by The BeeMD photo collection Queen on frame of ripening nectar above queen excluder; photo by The BeeMD photo collection
Queen in honey super
Queen on frame of ripening nectar above queen excluder; photo by The BeeMD photo collection
Adding queen excluder above brood before supering; photo by Dewey M. Caron Adding queen excluder above brood before supering; photo by Dewey M. Caron
Queen in honey super
Adding queen excluder above brood before supering; photo by Dewey M. Caron

Queen excluder; photo by The BeeMD photo collection Queen excluder; photo by The BeeMD photo collection
Queen in honey super
Queen excluder; photo by The BeeMD photo collection
Workers aggressively seeking to harm caged queen; photo by Dewey M. Caron Workers aggressively seeking to harm caged queen; photo by Dewey M. Caron
Queen rejection
Workers aggressively seeking to harm caged queen; photo by Dewey M. Caron
Adult mated queen introduction cage; photo by The BeeMD photo collection Adult mated queen introduction cage; photo by The BeeMD photo collection
Queen rejection
Adult mated queen introduction cage; photo by The BeeMD photo collection
Observe the behavior of workers toward confined queen at introduction; photo by Penn State University Observe the behavior of workers toward confined queen at introduction; photo by Penn State University
Queen rejection
Observe the behavior of workers toward confined queen at introduction; photo by Penn State University

Bees balling (rejecting) queen; photo by Wyatt Mangum Bees balling (rejecting) queen; photo by Wyatt Mangum
Queen rejection
Bees balling (rejecting) queen; photo by Wyatt Mangum
Small-bodied queen; could be due to poor genetics or queen shut down; photo by The BeeMD photo collection Small-bodied queen; could be due to poor genetics or queen shut down; photo by The BeeMD photo collection
Queen shut down
Small-bodied queen; could be due to poor genetics or queen shut down; photo by The BeeMD photo collection
Emergency queen cells; colony is queenless until a virgin hatches; photo by The BeeMD photo collection Emergency queen cells; colony is queenless until a virgin hatches; photo by The BeeMD photo collection
Queenless colony
Emergency queen cells; colony is queenless until a virgin hatches; photo by The BeeMD photo collection
Multiple eggs in worker cells; sign of a laying worker colony that is queenless; photo by The BeeMD photo collection Multiple eggs in worker cells; sign of a laying worker colony that is queenless; photo by The BeeMD photo collection
Queenless colony
Multiple eggs in worker cells; sign of a laying worker colony that is queenless; photo by The BeeMD photo collection

Queenless colony brood area filled with nectar in cells; photo by Steve Repasky Queenless colony brood area filled with nectar in cells; photo by Steve Repasky
Queenless colony
Queenless colony brood area filled with nectar in cells; photo by Steve Repasky
Drone-laying queen is not a queenless colony; photo by Dewey M. Caron Drone-laying queen is not a queenless colony; photo by Dewey M. Caron
Queenless colony
Drone-laying queen is not a queenless colony; photo by Dewey M. Caron
Two virgin queens fighting; photo by Beespoke.info Two virgin queens fighting; photo by Beespoke.info
Queens fighting
Two virgin queens fighting; photo by Beespoke.info
Dead queen; photo by Evan Fuller Dead queen; photo by Evan Fuller
Queens fighting
Dead queen; photo by Evan Fuller

Bees robbing hive at left; photo by Ana Heck Bees robbing hive at left; photo by Ana Heck
Robbing
Bees robbing hive at left; photo by Ana Heck
Robbing; photo by Honey Bee Health Coalition Robbing; photo by Honey Bee Health Coalition
Robbing
Robbing; photo by Honey Bee Health Coalition
Robbing screen at colony entrance; photo by Eric Mussen (deceased), UC Davis Robbing screen at colony entrance; photo by Eric Mussen (deceased), UC Davis
Robbing
Robbing screen at colony entrance; photo by Eric Mussen (deceased), UC Davis
Hive entrance protected with robbing screen; photo by Dewey M. Caron Hive entrance protected with robbing screen; photo by Dewey M. Caron
Robbing
Hive entrance protected with robbing screen; photo by Dewey M. Caron

Robbing by bees and yellowjackets at external Boardman feeder; photo by Dewey M. Caron Robbing by bees and yellowjackets at external Boardman feeder; photo by Dewey M. Caron
Robbing
Robbing by bees and yellowjackets at external Boardman feeder; photo by Dewey M. Caron
Dead prepupae with SBV removed from the cell; photo by Robert Snyder Dead prepupae with SBV removed from the cell; photo by Robert Snyder
Sacbrood virus
Dead prepupae with SBV removed from the cell; photo by Robert Snyder
Dead prepupae with sacbrood virus, removed from cell; photo by The BeeMD photo collection Dead prepupae with sacbrood virus, removed from cell; photo by The BeeMD photo collection
Sacbrood virus
Dead prepupae with sacbrood virus, removed from cell; photo by The BeeMD photo collection
Capped cells opened to view SBV-infected larvae; photo by Robert Snyder Capped cells opened to view SBV-infected larvae; photo by Robert Snyder
Sacbrood virus
Capped cells opened to view SBV-infected larvae; photo by Robert Snyder

SBV; photo by Robert Snyder SBV; photo by Robert Snyder
Sacbrood virus
SBV; photo by Robert Snyder
SBV; photo by Robyn Underwood SBV; photo by Robyn Underwood
Sacbrood virus
SBV; photo by Robyn Underwood
Scenting worker; photo by Robert Snyder Scenting worker; photo by Robert Snyder
Scenting
Scenting worker; photo by Robert Snyder
Scenting worker bee; photo by The BeeMD photo collection Scenting worker bee; photo by The BeeMD photo collection
Scenting
Scenting worker bee; photo by The BeeMD photo collection