Exotic
Cryptophlebia Walsingham (Tortricidae: Olethreutinae: Grapholitini)
The genus Cryptophlebia contains over 50 described species and is distributed in all regions except the Nearctic. Adults are moderately sized with brown forewings and variable, subdued patterns. Most species have a darker brown pretornal patch that is more pronounced in females. Males have a variety of sex scales on the legs, abdomen, and in special pockets on the hindwing. The male genitalia are characterized by swollen valvaevalva:
an appendage flanking the intromittent organ that is used to clasp the female during copulation
with multiple long spines on the inner surface of the cuculluscucullus:
the distal portion of the male valva
. The female genitalia are characterized by a ovate or V-shaped sterigmasterigma:
the sclerotized region surrounding the female ostium bursae 
, an anterioranterior:
before, to the front, toward the head 
widening of the ductus bursaeductus bursae:
a membranous tube connecting the ostium bursae to the corpus bursae
, and two signasignum:
a sclerotized projection or patch on the interior of the corpus bursae
in the corpus bursaecorpus bursae:
a dilated membranous sac at the anterior end of the bursa copulatrix
(Komai 1999Komai 1999:
Komai, F. 1999. A taxonomic review of the genus Grapholita and allied genera (Lepidoptera: Tortricidae) in the Palaearctic region. Entomologica Scandinavica Supplement 55. 226 pp.).
Cryptophlebia larvae are borers in seeds, nuts, fruits, pods, and stems. Larvae are whitish or pinkish with dark pinaculapinaculum:
flattened sclerotized plates on a caterpillar that bear the setae
and a brown head and prothoracic shieldprothoracic shield:
a sclerotized plate on the dorsal surface of the prothorax 
. An anal combanal comb:
a toothed structure on the last abdominal segment used to eject frass away from the feeding larva; also termed "anal fork" 
is absent in most species. The large prespiracular pinaculumpinaculum:
flattened sclerotized plates on a caterpillar that bear the setae on T1 extends under the spiracle and the spiracle on A8 is located on the posterodorsal margin of the segment; these two characters group Cryptophlebia larvae with other closely related genera such as Gymnandrosoma and Thaumatotibia.
This genus contains several important pest species. The three treated here, the koa seedworm (C. illepida), macadamia nut borer (C. ombrodelta), and litchi moth (C. peltastica ), are important pests of litchi and macadamia. Cryptophlebia larvae are intercepted very frequently at U.S. ports of entry, although it is likely that these are not distinguished from larvae of related species, such as false codling moth (Thaumatotibia leucotreta), which was previously placed in Cryptophlebia.
Exotic species
