Coelostathma discopunctana

Type

Native

Taxonomy

Coelostathma discopunctana Clemens (Tortricidae: Tortricinae: Sparganothini)

Common name: batman moth

Synonyms: discipunctana (Coelostathma)

Adult Recognition

FWLFWL:
forewing length; the distance from the base of the forewing to the apex, including fringe
: 6.5-6.8 mm (males); 6.9-7.4 mm (females)

Head and thorax pale yellow to pale brown; labial palpilabial palpi:
a pair of sensory appendages that project from the lower part of the head; usually covered in scales and three-segmented
long; forewing with broad, somewhat diffuse median fasciafascia:
a dark transverse band on the forewing
, darker along inner margininner margin:
see dorsum
in females; a small black dot present near the apexapex:
the point furthest removed from the base or at the end of the costal area
of the discal cell; faint reticulationsreticulations:
covered with a network of lines or mesh; in reference to wing pattern, usually thin horizontal lines
present over much of forewing; male forewing costal foldforewing costal fold:
a flap or fold at the base of the forewing that contains specialized sex scales
absent; female forewing similar to male but slightly darker, markings less distinct. hindwing in both sexes white to pale brown.

Male genitalia are characterized by a long, slender, curved uncusuncus:
a sclerotized process which is fused to the posterodorsal margin of tergum IX
; somewhat triangular valvaevalva:
an appendage flanking the intromittent organ that is used to clasp the female during copulation
; and a well-defined sacculussacculus:
the ventral margin of the male valva
with a slender anterioranterior:
before, to the front, toward the head
process extending into the disc of the valvavalva:
an appendage flanking the intromittent organ that is used to clasp the female during copulation
. The female genitalia are characterized by a weakly bilobed sterigmasterigma:
the sclerotized region surrounding the female ostium bursae
; signumsignum:
a sclerotized projection or patch on the interior of the corpus bursae
present as a broad, curved band in the caudal region of the corpus bursaecorpus bursae:
a dilated membranous sac at the anterior end of the bursa copulatrix
.

Larval Morphology

The following account is summarized from Powell & Brown (2012).

Mature larva 12-14 mm in length. Head amber, with black pigmentation laterally; body translucent in early instars, becoming pale green-gray with age, prothoracic shieldprothoracic shield:
a sclerotized plate on the dorsal surface of the prothorax
concolorous with broad, irregularly triangular patch of darker pigmentation laterally; semitransparent; pinaculapinaculum:
flattened sclerotized plates on a caterpillar that bear the setae
 moderately small, darkening with age; anal fork well-developed. SV group on A1, 2, 7, 8, 9 usually 3:3:3:2:2.

A complete, detailed description of larval chaetotaxychaetotaxy:
the arrangement of setae (in reference to Lepidoptera larvae), often depicted on a "setal map"
is available in Powell & Brown (2012).

Similar Species

Adults of C. discopunctana could be confused with Sparganothoides lentiginosana (Walsingham), a similar-looking southeastern U.S. species. However, adults of S. lentiginosana lack the defined median fasciafascia:
a dark transverse band on the forewing
, the distinct discal spot, and abdominal dorsaldorsal:
upper, to the top, on the back
pits found in both sexes of C. discopunctana. Males of S. lentiginosana possess a forewing costal foldforewing costal fold:
a flap or fold at the base of the forewing that contains specialized sex scales
, which males of C. discopunctana lack.

Biology

The following account is summarized from unpublished notes by Busck included in Powell & Brown (2012).

In Virginia, females of C. discopunctana lay small patches of 5-20 eggs over about a one-week period in September on leaves of the host plant. Larvae hatch about one week later and feed by constructing an open silken chamber on the host. Development takes between 45-50 days, but probably fewer in the summer months and further south. Overwintering occurs as late instar larvae or as pupae. At least two, possibly more, generations occur per year. The first generation of adults peaks in late May into June and the second from late July into August and early September. Most adult records come from May through September, but adults may be active year-round along the Gulf Coast.

Larvae have been reared from a few different low-growing herbs as well as apple.

Host plant Host plant family Reference(s)
Impatiens sp. Balsaminaceae USNM collectionUSNM collection:
USNM collection. Based on identified reared specimens in the collection of the National Museum of Natural History, Washington, D.C.
; Powell & Brown 2012
Desmodium sp. Fabaceae Powell 2006Powell 2006:
Powell, J. A. 2006. Database of Lepidoptera rearing lots, 1960-2005. University of California Berkeley, CA.
Trifolium sp. Fabaceae Fernald 1882aFernald 1882a:
Fernald, C. H. 1882a. A synonymical catalogue of the described Tortricidae of North America, north of Mexico. Transactions of the American Entomological Society. 10: 1-64.
; Forbes 1923Forbes 1923:
Forbes, W. M. T. 1923. The Lepidoptera of New York and neighboring states. Primitive forms, Microlepidoptera, Pyraloids, Bombyces. Cornell Univ. Argic. Exp. Sta., Mem. 68. 729 pp.
Habenaria sp. Orchidaceae USNM collectionUSNM collection:
USNM collection. Based on identified reared specimens in the collection of the National Museum of Natural History, Washington, D.C.
; Powell & Brown 2012
Fragaria x ananassa Rosaceae USNM collectionUSNM collection:
USNM collection. Based on identified reared specimens in the collection of the National Museum of Natural History, Washington, D.C.
; Powell & Brown 2012
Malus domestica Rosaceae USNM collectionUSNM collection:
USNM collection. Based on identified reared specimens in the collection of the National Museum of Natural History, Washington, D.C.

Host plant table (embedded)

View full screen host table here

Distribution

Coelostathma discopunctana is broadly distributed in eastern North America, from Nova Scotia and northern Minnesota south to Florida and eastern Texas.

Links

Additional photos and a distribution map of this species in North America are available at Moth Photographers Group.
 Male. © John W. Brown. Image used with permission.
Male. © John W. Brown. Image used with permission.
 Female. © John W. Brown. Image used with permission.
Female. © John W. Brown. Image used with permission.