Native
Clepsis persicana (Fitch) (Tortricidae: Tortricinae: Archipini)
Common names: white triangle tortrix, green needleworm
Synonyms: blandana (Ditula), conigerana (Tortrix), fragariana (Lozotaenia)
Subspecies: C. persicana forbesi (British Columbia)
FWLFWL:
forewing length; the distance from the base of the forewing to the apex, including fringe
: 8.5-10.5 mm (male); 10.0-11.0 mm (female)
Forewings are orange basally, becoming darker and purplish towards the pale termentermen:
the outer edge of the forewing
. Most individuals have a well-defined white costal triangle, although this marking is reduced in some western phenotypes. Males have a long forewing costal foldforewing costal fold:
a flap or fold at the base of the forewing that contains specialized sex scales
that extends to nearly half the length of the costacosta:
the anterior margin of each wing
.
Male genitalia are characterized by a large, bulbous uncusuncus:
a sclerotized process which is fused to the posterodorsal margin of tergum IX
; an incomplete, spined uncusuncus:
a sclerotized process which is fused to the posterodorsal margin of tergum IX
; and small, membranous valvaevalva:
an appendage flanking the intromittent organ that is used to clasp the female during copulation
. Female genitalia are characterized by a long ductus bursaeductus bursae:
a membranous tube connecting the ostium bursae to the corpus bursae
without a cestumcestum:
a long, bandlike sclerotization of the wall of the ductus bursae
; and a small, daggar-like signumsignum:
a sclerotized projection or patch on the interior of the corpus bursae
in the corpus bursaecorpus bursae:
a dilated membranous sac at the anterior end of the bursa copulatrix
with a robust capitulumcapitulum:
in reference to the female signum; a knoblike projection
.
The following account is summarized from MacKay (1962a).
Mature larvae approximately 14-18 mm in length; width of head averaging 1.1-1.2 mm; head, thoracic shield brownish yellow; body pale green; anal fork well-developed; SV group on A1, 2, 7, 8, 9 numbering 3:3:3:2:2.
Detailed figures of larval chaetotaxychaetotaxy:
the arrangement of setae (in reference to Lepidoptera larvae), often depicted on a "setal map"
are available in MacKay (1962a).
Acleris holmiana is superficially similar to Clepsis persicana, but can be distinguished by the absence of a male forewing costal foldforewing costal fold:
a flap or fold at the base of the forewing that contains specialized sex scales
. With this lone exception, forewing pattern is sufficient to distinguish adults of C. persicana from all other Nearctic Tortricidae.
Larvae of Clepsis persicana may be confused with a number of other tortricid larvae, including those of Clepsis, Argyrotaenia, Choristoneura, and Epiphyas postvittana.
The following account is summarized from Powell (1964b) and Chapman & Lienk (1971).
Clepsis persicana completes a single generation per year. Adults are present from June to August. Eggs are laid in overlapping patches on the upper surface of leaves or on smooth bark. Newly hatched larvae are incapable of constructing their own shelters and lower themselves on a silk thread searching for abandonded shelters of other tortricid species, such as Spilonota ocellana. If a suitable shelter is not found, larvae will descend to the ground and feed on cover plants for the remainder of the summer and autumn. Mid-instar larvae overwinter and complete development in the spring, where they may ascend trees or other woody plants. Larvae can cause damage to fruit by webbing leaves to the fruit or feeding in the calyx. Pupation occurs under bark or in fallen leaves near the base of a tree.
Larvae are generalists and have been recorded from over 40 species of deciduous and coniferous trees. Although Fitch described C. persicana from larvae he reared on peach (Prunus persica), this plant is not thought to be a primary host.
Host plant | Host plant family | Reference(s) |
Acer negundo | Aceraceae | Prentice 1966Prentice 1966: Prentice, R. M. 1966. Vol. 4. Microlepidoptera. In : Forest Lepidoptera of Canada recorded by the Forest Insect Survey. Dept. For. Canada Publ. 1142: 543-840. |
Acer sp. | Aceraceae | MacKay 1962aMacKay 1962a: MacKay, M. R. 1962a. Larvae of the North American Tortricinae (Lepidoptera: Tortricidae). Canadian Entomologist, Supplement 28: 1-182., Powell 1964bPowell 1964b: Powell, J. A. 1964b. Biological and taxonomic studies on tortricine moths, with reference to the species in California. University of California Publications in Entomology. Vol. 32. 317 pp. |
Osmorhiza berteroi | Apiaceae | Powell 1964bPowell 1964b: Powell, J. A. 1964b. Biological and taxonomic studies on tortricine moths, with reference to the species in California. University of California Publications in Entomology. Vol. 32. 317 pp. |
Solidago sp. | Asteraceae | Schaffner 1959Schaffner 1959: Schaffner, J. V. 1959. Microlepidoptera and their parasites reared from field collections in the northeastern United States. USDA, Misc. Publ. 767. 97 pp.; Powell 1964bPowell 1964b: Powell, J. A. 1964b. Biological and taxonomic studies on tortricine moths, with reference to the species in California. University of California Publications in Entomology. Vol. 32. 317 pp. |
Alnus incana | Betulaceae | Prentice 1966Prentice 1966: Prentice, R. M. 1966. Vol. 4. Microlepidoptera. In : Forest Lepidoptera of Canada recorded by the Forest Insect Survey. Dept. For. Canada Publ. 1142: 543-840. |
Alnus viridis crispa | Betulaceae | Prentice 1966Prentice 1966: Prentice, R. M. 1966. Vol. 4. Microlepidoptera. In : Forest Lepidoptera of Canada recorded by the Forest Insect Survey. Dept. For. Canada Publ. 1142: 543-840.; Ferguson 1975Ferguson 1975: Ferguson, D. C. 1975. Host records for Lepidoptera reared in eastern North America. United States Department of Agriculture Technical Bulletin 1521. 49 pp. |
Alnus sp. | Betulaceae | Ferguson 1975Ferguson 1975: Ferguson, D. C. 1975. Host records for Lepidoptera reared in eastern North America. United States Department of Agriculture Technical Bulletin 1521. 49 pp. |
Betula nana | Betulaceae | Prentice 1966Prentice 1966: Prentice, R. M. 1966. Vol. 4. Microlepidoptera. In : Forest Lepidoptera of Canada recorded by the Forest Insect Survey. Dept. For. Canada Publ. 1142: 543-840. |
Betula papyrifera | Betulaceae | Prentice 1966Prentice 1966: Prentice, R. M. 1966. Vol. 4. Microlepidoptera. In : Forest Lepidoptera of Canada recorded by the Forest Insect Survey. Dept. For. Canada Publ. 1142: 543-840. |
Betula sp. | Betulaceae | Ferguson 1975Ferguson 1975: Ferguson, D. C. 1975. Host records for Lepidoptera reared in eastern North America. United States Department of Agriculture Technical Bulletin 1521. 49 pp. |
Corylus sp. | Betulaceae | Prentice 1966Prentice 1966: Prentice, R. M. 1966. Vol. 4. Microlepidoptera. In : Forest Lepidoptera of Canada recorded by the Forest Insect Survey. Dept. For. Canada Publ. 1142: 543-840. |
Cornus canadensis | Cornaceae | Ferguson 1975Ferguson 1975: Ferguson, D. C. 1975. Host records for Lepidoptera reared in eastern North America. United States Department of Agriculture Technical Bulletin 1521. 49 pp. |
Rhododendron canadense | Ericaceae | Ferguson 1975Ferguson 1975: Ferguson, D. C. 1975. Host records for Lepidoptera reared in eastern North America. United States Department of Agriculture Technical Bulletin 1521. 49 pp. |
Vaccinium sp. | Ericaceae | LACM IndexLACM Index: LACM Index. Records from the card file at the Los Angeles County Museum of Natural History, Los Angeles, California; transcribed by Gaeden Robinson (BMNH).; Ferguson 1975Ferguson 1975: Ferguson, D. C. 1975. Host records for Lepidoptera reared in eastern North America. United States Department of Agriculture Technical Bulletin 1521. 49 pp. |
Frasera fastigiata | Gentianaceae | Powell 1964bPowell 1964b: Powell, J. A. 1964b. Biological and taxonomic studies on tortricine moths, with reference to the species in California. University of California Publications in Entomology. Vol. 32. 317 pp. |
Frasera sp. | Gentianaceae | Powell 1964bPowell 1964b: Powell, J. A. 1964b. Biological and taxonomic studies on tortricine moths, with reference to the species in California. University of California Publications in Entomology. Vol. 32. 317 pp. |
Ribes sp. | Grossulariaceae | Prentice 1966Prentice 1966: Prentice, R. M. 1966. Vol. 4. Microlepidoptera. In : Forest Lepidoptera of Canada recorded by the Forest Insect Survey. Dept. For. Canada Publ. 1142: 543-840. |
Maianthemum canadense | Liliaceae | Ferguson 1975Ferguson 1975: Ferguson, D. C. 1975. Host records for Lepidoptera reared in eastern North America. United States Department of Agriculture Technical Bulletin 1521. 49 pp. |
Comptonia peregrina | Myrtaceae | Ferguson 1975Ferguson 1975: Ferguson, D. C. 1975. Host records for Lepidoptera reared in eastern North America. United States Department of Agriculture Technical Bulletin 1521. 49 pp. |
Fraxinus sp. | Oleaceae | Powell 1964bPowell 1964b: Powell, J. A. 1964b. Biological and taxonomic studies on tortricine moths, with reference to the species in California. University of California Publications in Entomology. Vol. 32. 317 pp. |
Abies balsamea | Pinaceae | Prentice 1966Prentice 1966: Prentice, R. M. 1966. Vol. 4. Microlepidoptera. In : Forest Lepidoptera of Canada recorded by the Forest Insect Survey. Dept. For. Canada Publ. 1142: 543-840. |
Abies concolor |