Neospondylis

Subfamily

Spondylidinae

Diagnosis - adult

Body length: 8–20mm. 
Eyes: eye interommatidial setaeseta:
a sclerotized hair-like projection of the cuticle
absent, eye entire/shallowly emarginateemarginate:
notched at the margin
, eye ommatidial density fine. 
AntennaeAntenna:
in larval and adult insects, paired segmented appendages, borne one on each side of the head, functioning as sense organs and bearing a large number of sensilla
: antennal length barely surpassing pronotumpronotum:
the upper and dorsal part of the prothorax
or shorter, antennal flagellar segments elongateelongate:
much longer than wide
, rarely quadrate, scapescape:
the first proximal segment of the antenna
smooth/punctate at apexapex:
end of any structure distad to the base
, antennal scapescape:
the first proximal segment of the antenna
≥ segment 3. 
Pronotumpronotum:
the upper and dorsal part of the prothorax
: pronotumpronotum:
the upper and dorsal part of the prothorax
shape transversetransverse:
broader than long
, pronotumpronotum:
the upper and dorsal part of the prothorax
lateral armature absent. 
Prosternum: prosternal process dilated at apexapex:
end of any structure distad to the base
, rarely not dilated at apexapex:
end of any structure distad to the base
, procoxal cavities closed posteriorly. 
ElytraElytron:
the leathery forewing of beetles, serving as a covering for the hind wings, commonly meeting opposite elytron in a straight line down the middle of the dorsum in repose
: elytral length reaching or close to end of abdomen, elytral apicesapex:
end of any structure distad to the base
rounded or truncatetruncate:
cut off squarely at the tip
, elytral color black, brown, or reddish, elytral color pattern absent. 
Legs: visible tarsomeres: 5, femora slender or robust, protibial spurs: 2, tarsal claws simple. 

Head large, almost as broad as pronotumpronotum:
the upper and dorsal part of the prothorax
; mandibles shorter than head, cutting edge with a basal tooth; second segment of maxillary palpi slightly shorter than third; galea transversetransverse:
broader than long
, apically expanded, spatuliform; lacinia vestigial, ligula attenuateattenuate:
gradually tapering apically
, not bilobed apically; antennaeantenna:
in larval and adult insects, paired segmented appendages, borne one on each side of the head, functioning as sense organs and bearing a large number of sensilla
inserted in front of eyes near the basebase:
the part of any appendage or structure that is nearest the body
of pronotumpronotum:
the upper and dorsal part of the prothorax
, not or hardly attaining the basebase:
the part of any appendage or structure that is nearest the body
of elytraelytron:
the leathery forewing of beetles, serving as a covering for the hind wings, commonly meeting opposite elytron in a straight line down the middle of the dorsum in repose
, third segment about twice as long as second, third and fourth segments globose, the following moderately flattened; segments fifth to tenth with two separate small sensory areas at each side of their apexapex:
end of any structure distad to the base
. Pronotumpronotum:
the upper and dorsal part of the prothorax
cordiform, narrowing posteriorly from about apical 1/3. Prosternum distinctly convex in front, with process parallel-sided, strongly expanded apically, coxal cavities strongly angulate laterally, closed behind (pleural and prosternal processes connected but not fused); mesothoracic process not broader than the prosternal one, moderately expanded behind, bilobed at apexapex:
end of any structure distad to the base
; coxal cavities open externally to the epimera; metathoracic episterna very narrow. Each elytronelytron:
the leathery forewing of beetles, serving as a covering for the hind wings, commonly meeting opposite elytron in a straight line down the middle of the dorsum in repose
with 2-3 more or less distinct glabrousglabrous:
smooth, devoid of pubescence; devoid of any sculpturing
costae, sometimes obsolete, rarely absent. Legs short, with hind femora very robust, enlarged; tibiaetibia:
the leg segment distal to the femur, proximal to the tarsus
granulated or finely spined on the outer side; front tibiaetibia:
the leg segment distal to the femur, proximal to the tarsus
densely punctatepunctate:
set with fine, impressed points or punctures appearing as pin-pricks
externally; hind tibiaetibia:
the leg segment distal to the femur, proximal to the tarsus
without a distinct lamella at the outer apexapex:
end of any structure distad to the base
and with two long apical spines (adapted from Sama 2005Sama 2005:
Sama G. 2005. Description of Neospondylis gen. nov. from North America and Mexico (Coleoptera, Cerambycidae, Spondylinae). Les Cahiers Magellanes 43: 1-10, 38 figs.
).

Similar genera

Spondylis

Differential diagnosis

The smooth cutting edge of mandibles, closed coxal cavities, non-globose pronotumpronotum:
the upper and dorsal part of the prothorax
, and lack of teeth on the outer edge of tibiaetibia:
the leg segment distal to the femur, proximal to the tarsus
will distinguish Neospondylis from Spondylis

Spondylis Fabricius differs from Neospondylis as follows: mandibles with small teeth on the cutting edge; second segment of maxillary palpi shorter than third; galea small, digitiform, ligula strongly expanded laterally, deeply bilobed, with or without a prominent median tooth at apexapex:
end of any structure distad to the base
. Pronotumpronotum:
the upper and dorsal part of the prothorax
globose, rounded at sides, shortly parallel-sided before the basebase:
the part of any appendage or structure that is nearest the body
. AntennaeAntenna:
in larval and adult insects, paired segmented appendages, borne one on each side of the head, functioning as sense organs and bearing a large number of sensilla
with third segment longer than fourth; segments third to tenth flattened above and with a wide sensory area on the outer side. Prosternum moderately convex in front, with process narrower, parallel sided, slightly expanded apically, coxal cavities open behind (pleural and prosternal processes not connected); mesothoraxmesothorax:
the second or middle thoracic segment bearing the midlegs and the forewings
with process evidently broader than the prosternal one, parallel-sided, not expanded behind, bilobed at apexapex:
end of any structure distad to the base
, episterna distinctly wider. TibiaeTibia:
the leg segment distal to the femur, proximal to the tarsus
on the outer side with several prominent teeth; front tibiaetibia:
the leg segment distal to the femur, proximal to the tarsus
flattened and shining externally; all tibiaetibia:
the leg segment distal to the femur, proximal to the tarsus
with a larger conspicuous lamella on the outer apexapex:
end of any structure distad to the base
(Sama 2005Sama 2005:
Sama G. 2005. Description of Neospondylis gen. nov. from North America and Mexico (Coleoptera, Cerambycidae, Spondylinae). Les Cahiers Magellanes 43: 1-10, 38 figs.
).

Distribution

Nearctic: coniferous forests of North America from Alaska southeast to the Great Lakes region, south into the Rocky Mountains, and along the western coast; Mexico

Recorded host plants

Picea, Pinus

Remarks

2 species

Taxonomy

Neospondylis Sama, 2005Sama, 2005:
Sama G. 2005. Description of Neospondylis gen. nov. from North America and Mexico (Coleoptera, Cerambycidae, Spondylinae). Les Cahiers Magellanes 43: 1-10, 38 figs.
, Les Cahiers Magellanes 43: 2.

  Neospondylis upiformis  dorsal

Neospondylis upiformis dorsal

  Neospondylis upiformis  ventral

Neospondylis upiformis ventral

  Neospondylis upiformis  frontal

Neospondylis upiformis frontal

  Neospondylis upiformis  lateral

Neospondylis upiformis lateral

 EDRR5a  Neospondylis upiformis  voucher
EDRR5a Neospondylis upiformis voucher