Gallery

Onthophagus tuberculifrons female head; photo by E.L. Engasser

distribution map for Onthophagus tuberculifrons

Oryctes boas male; photo by E.L. Engasser

Oryctes boas male; photo by E.L. Engasser

Oryctes centaurus male; photo by E.L. Engasser

Oryctes centaurus male; photo by E.L. Engasser

Oryctes nasicornis male; photo by E.L. Engasser

Oryctes nasicornis male; photo by E.L. Engasser

Oryctes nasicornis female; photo by E.L. Engasser

Oryctes nasicornis female; photo by E.L. Engasser

Oryctes rhinoceros characteristic damage to palm trees; photo by M.L. Jameson

Oryctes rhinoceros characteristic damage to palm trees; photo by M.L. Jameson

distribution map for Orcytes rhinoceros

Oryctes rhinoceros male; photo by E.L. Engasser

Oryctes rhinoceros male; photo by E.L. Engasser

Oryctes rhinoceros male; photo by E.L. Engasser

Oryctes rhinoceros male; photo by E.L. Engasser

Oryctes rhinoceros female; photo by E.L. Engasser

Oryctes rhinoceros female; photo by E.L. Engasser

Oryctes rhinoceros male pygidium, lateral view; photo by E.L. Engasser

Oryctes rhinoceros female pygidium, lateral view; photo by E.L. Engasser

Oryctes rhinoceros male genitalia, caudal view; photo by E.L. Engasser

Oryctes rhinoceros male genitalia, lateral view; photo by E.L. Engasser

adult Oryctes rhinoceros entangled in net trap in Guam; photo by M.L. Jameson

newly emerged Oryctes rhinoceros surrounded by pupae; photo by E.L. Engasser

Oryctes rhinoceros (coconut rhinoceros beetle) larva in comparison with Protaetia orientalis (Asian flower beetle); photo by M.L. Jameson

coconut palm showing characteristic Oryctes damage in Hawaii; photo by M.L. Jameson

coconut palm showing characteristic Oryctes damage to fronds in Guam; photo by M.L. Jameson

distribution map for Orcytes rhinoceros

Papuana uninodis male; photo by E.L. Engasser

Papuana uninodis male; photo by E.L. Engasser

Papuana lansbergei male; photo by E.L. Engasser