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Morphological Structure

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Phytophthora hedraiandra

Phytophthora hedriandra (CPHST BL 4) sexual phase of the ex-type: smooth oogonium with paragynous antheridium and aplerotic oospore; photo by Gloria Abad, USDA-APHIS-PPQ.

Phytophthora hedraiandra

Phytophthora hedriandra (CPHST BL 4) asexual phase of the ex-type: sporangia and oospores; photo by Gloria Abad USDA-APHIS-PPQ.

Phytophthora hedraiandra

Phytophthora hedriandra (CPHST BL 4) asexual phase of the ex-type: sporangium papillate with caducous short pedicel; photos by Gloria Abad USDA-APHIS-PPQ.

Phytophthora heterospora
Phytophthora spp. in Clade 4: portion of the seven-loci ML phylogeny featuring the type cultures of 212 described species (by T. Bourret). Notice the position of P. heterospora Ex-type CBS 148034 = S&T BL 119. Gloria Abad, USDA S&T.

Phytophthora heterospora
Phytophthora spp. in Clade 4: Morphological Tabular key (PDF) and Tabular key legends (PDF) in IDphy2 KEY SECTION. Notice the data of P. heterospora Ex-type CBS 148034 = S&T BL 119. Gloria Abad, USDA S&T.
Phytophthora heterospora

colony morphology after 4 d growth at 20 ºC in the dark on carrot agar, V8A, CA, MEA, and PDA (from left to right)

Phytophthora heterospora

laterally attached papillate sporangium; scale bar = 25µm

Phytophthora heterospora

oogonia globose with aplerotic oospores, short tapering bases, often with aborted oospore; antheridia bicellular and often with finger-like projection (from left to right); scale bar = 25 μm


Phytophthora heterospora

ovoid pseudoconidia forming monochasial helicoid sympodia, still attached to the conidiophores or recently detached with nodose swellings at the insertion points of the pseudoconidia to the conidiophores; spherical to ovoid pseudoconidia with short pre-formed pedicels; ovoid, laterally attached papillate sporangium; pseudoconidia germinating by forming papillate microsporangia and a new conidium (from left to right); scale bar = 25µm

Phytophthora heterospora

disease symptoms on Olea europaea: 4-year-old tree showing chlorosis and increased transparency of the crown; orange-brown and flame shaped lesions on the stem; single-spot bark lesions on the main stem

Phytophthora heveae
Phytophthora spp. in Clade 5: portion of the seven-loci ML phylogeny featuring the type cultures of 212 described species (by T. Bourret). Notice the position of P. heveae Ex-type CBS 296.29 = S&T BL 67. Gloria Abad, USDA S&T.
Phytophthora heveae
Phytophthora spp. in Clade 5: Morphological Tabular key (PDF) and Tabular key legends (PDF) in IDphy2 KEY SECTION. Notice the data of P. heveae Ex-type CBS 296.29 = S&T BL 67. Gloria Abad, USDA S&T.

Phytophthora heveae

Phytophthora heveae (CPHST BL 67) colonies of the ex-type grown for 7 days on (a) V8® Agar, (b) potato dextrose agar, and (c) malt extract agar; photo by Krysta Jennings and Leandra Knight, USDA-APHIS-PPQ

Phytophthora heveae

Phytophthora heveae (P10167, selected specimen) asexual phase: (a) sporangia in sporangiophores; (b–g) papillate sporangia, persistent and caducous - with pedicels (blue arrows); (f) bipapillate sporangium, (g) sporangium with external proliferation; photos by Gloria Abad, USDA-APHIS-PPQ.

Phytophthora heveae

Phytophthora heveae (P10167, selected specimen) homothallic sexual phase: (a–d) smooth-walled oogonia frequently with tapered bases; amphigynous antheridia and aplerotic oospores; photos by Gloria Abad, USDA-APHIS-PPQ.

Phytophthora heveae

Phytophthora heveae (P10167, selected specimen) asexual phase: persistent papillate sporangia and sporangiophores; photo by Gloria Abad, USDA-APHIS-PPQ.


Phytophthora heveae

Phytophthora heveae (P10167, selected specimen) asexual phase: papillate sporangium with persistent pedicel; photo by Gloria Abad, USDA-APHIS-PPQ.

Phytophthora heveae

Phytophthora heveae (P10167, selected specimen) asexual phase: papillate sporangium; photo by Gloria Abad, USDA-APHIS-PPQ.

Phytophthora heveae

Phytophthora heveae (P10167, selected specimen) asexual phase: papillate sporangium; photo by Gloria Abad, USDA-APHIS-PPQ.

Phytophthora heveae

Phytophthora heveae (P10167, selected specimen) asexual phase: bipapillate caducous sporangium with pedicel; photo by Gloria Abad, USDA-APHIS-PPQ.


Phytophthora heveae

Phytophthora heveae (P10167, selected specimen) asexual phase: papillate caducous sporangium with pedicel; photo by Gloria Abad, USDA-APHIS-PPQ.

Phytophthora heveae

Phytophthora heveae (P10167, selected specimen) asexual phase: sporangium with external proliferation; photo by Gloria Abad, USDA-APHIS-PPQ.

Phytophthora heveae

Phytophthora heveae (P10167, selected specimen) homothallic sexual phase: smooth-walled oogonium with tapered base; amphigynous antheridium and aplerotic oospore; photo by Gloria Abad, USDA-APHIS-PPQ.

Phytophthora heveae

Phytophthora heveae (P10167, selected specimen) homothallic sexual phase: smooth-walled oogonium with tapered base; amphigynous antheridium and aplerotic oospore; photo by Gloria Abad, USDA-APHIS-PPQ.


Phytophthora heveae

Phytophthora heveae (P10167, selected specimen) homothallic sexual phase: smooth-walled oogonia with tapered bases; amphigynous antheridia and aplerotic oospores; photo by Gloria Abad, USDA-APHIS-PPQ.

Phytophthora heveae

Phytophthora heveae (P10167, selected specimen) homothallic sexual phase: smooth-walled oogonia with tapered bases; amphigynous antheridia and aplerotic oospores; photo by Gloria Abad, USDA-APHIS-PPQ.

Phytophthora heveae

Phytophthora heveae (P10167, selected specimen) homothallic sexual phase: smooth-walled oogonia with tapered bases; amphigynous antheridia and aplerotic oospores; photo by Gloria Abad, USDA-APHIS-PPQ.

Phytophthora hibernalis (in progress - Abad et al. 2023b)
Phytophthora spp. in subclade 8c: portion of the seven-loci ML phylogeny featuring the type cultures of 212 described species (by T. Bourret). Notice the position of P. hibernalis selected specimen CBS 114104 = S&T BL 41G. Gloria Abad, USDA S&T.

Phytophthora hibernalis (in progress - Abad et al. 2023b)
Phytophthora spp. in subclade 8c: Morphological Tabular key (PDF) and Tabular key legends (PDF) in IDphy2 KEY SECTION. Notice the data of P. hibernalis selected specimen CBS 114104 = S&T BL 41G. Gloria Abad, USDA S&T.
Phytophthora hibernalis (in progress - Abad et al. 2023b)

Phytophthora hibernalis (CPHST BL 41G) colonies of the ex-epitype grown for 7 days on (a) V8® Agar, (b) potato dextrose agar, and (c) malt extract agar; photo by Krysta Jennings and Leandra Knight, USDA-APHIS-PPQ

Phytophthora hibernalis (in progress - Abad et al. 2023b)

Phytophthora hibernalis (CPHST BL 41G) asexual phase: (a–d) semipapillate sporangia, caducous with medium to long pedicels; photos by Gloria Abad, USDA-APHIS-PPQ.