PLANT FAMILY PLANT SPECIES PLANT COMMON NAME ASSOCIATED PEST REFERENCE LINK
Pinaceae Pinus aristata Rocky Mountain bristlecone pine Eucopina bobana Powell 1968

Powell 1968


Powell, J. A. 1968. Host associations and taxonomy of nearctic conifer cone moths in the genus Eucosma (Lepidoptera: Tortricidae). Hilgardia. 39: 1-36.
Eucopina spp.
Pinaceae Pinus cembroides Mexican pinyon pine Eucopina bobana Cibrian-Tovar et al. 1986

Cibrian-Tovar et al. 1986


Cibrian-Tovar, D., B. H. Ebel, H. O. Yates & J. T. Mendez-Montiel (eds.). 1986. Cone and seed insects of the Mexican conifers. Southeastern Forest Experiment Station, U.S. Department of Agriculture.
Eucopina spp.
Pinaceae Pinus contorta lodgepole pine Eucopina bobana Prentice 1966

Prentice 1966


Prentice, R. M. 1966. Vol. 4. Microlepidoptera. In: Forest Lepidoptera of Canada recorded by the Forest Insect Survey. Dept. For. Canada Publ. 1142: 543-840.
Eucopina spp.
Pinaceae Pinus edulis pinyon pine Eucopina bobana Forcella 1978

Forcella 1978


Forcella, F. 1978. Irregularity of pinyon cone production and its relation to pinyon cone moth predation. Madrono. 25: 170-172.
; Powell 1968

Powell 1968


Powell, J. A. 1968. Host associations and taxonomy of nearctic conifer cone moths in the genus Eucosma (Lepidoptera: Tortricidae). Hilgardia. 39: 1-36.
Eucopina spp.
Pinaceae Pinus flexilis limber pine Eucopina bobana Powell 1968

Powell 1968


Powell, J. A. 1968. Host associations and taxonomy of nearctic conifer cone moths in the genus Eucosma (Lepidoptera: Tortricidae). Hilgardia. 39: 1-36.
Eucopina spp.
Pinaceae Pinus jefferyi Jeffrey pine Eucopina bobana Heinrich 1923b

Heinrich 1923b


Heinrich, C. 1923b. Revision of the North American moths of the subfamily Eucosminae of the family Olethreutidae. Bulletin of the United States National Museum. 123: 1-298.
Eucopina spp.
Pinaceae Pinus monophylla single-leaf pinyon Eucopina bobana Forcella 1978, 1979

Forcella 1978, 1979



Forcella, F. 1978. Irregularity of pinyon cone production and its relation to pinyon cone moth predation. Madrono. 25: 170-172.


Forcella, F. 1979. Mast fruiting in trees: why and how in cembroid pines. Ph.D. dissertation, University of Oklahoma, Norman. 


; Powell 1968

Powell 1968


Powell, J. A. 1968. Host associations and taxonomy of nearctic conifer cone moths in the genus Eucosma (Lepidoptera: Tortricidae). Hilgardia. 39: 1-36.
Eucopina spp.
Pinaceae Pinus monophylla single-leaf pinyon Eucopina bobana Powell 2006

Powell 2006


Powell, J. A. 2006. Database of Lepidoptera rearing lots, 1960-2005. University of California Berkeley, CA.
Eucopina spp.
Pinaceae Pinus monticola western white pine Eucopina bobana Prentice 1966

Prentice 1966


Prentice, R. M. 1966. Vol. 4. Microlepidoptera. In: Forest Lepidoptera of Canada recorded by the Forest Insect Survey. Dept. For. Canada Publ. 1142: 543-840.
Eucopina spp.
Pinaceae Pinus pinceana weeping pinyon pine Eucopina bobana Cibrian-Tovar et al. 1986

Cibrian-Tovar et al. 1986


Cibrian-Tovar, D., B. H. Ebel, H. O. Yates & J. T. Mendez-Montiel (eds.). 1986. Cone and seed insects of the Mexican conifers. Southeastern Forest Experiment Station, U.S. Department of Agriculture.
Eucopina spp.
Pinaceae Pinus ponderosa Ponderosa pine Eucopina bobana Heinrich 1923b

Heinrich 1923b


Heinrich, C. 1923b. Revision of the North American moths of the subfamily Eucosminae of the family Olethreutidae. Bulletin of the United States National Museum. 123: 1-298.
Eucopina spp.
Pinaceae Pinus taeda loblolly pine Eucopina cocana Powell 1968

Powell 1968


Powell, J. A. 1968. Host associations and taxonomy of nearctic conifer cone moths in the genus Eucosma (Lepidoptera: Tortricidae). Hilgardia. 39: 1-36.
; Ebel et al. 1975

Ebel et al. 1975


Ebel, B.H., Flavell, T.H., Drake, L.E., Yates, H.O., DeBarr, G.L. 1975. Seed and cone insects of southern pines. General technical Report/SE-8, USDA Forest Service, Southern Forest Experiment Station. 39 pp.
Eucopina spp.
Pinaceae Picea sp. spruce Eucopina gloriola DeBoo et al. 1971

DeBoo et al. 1971


DeBoo, R. F., W. L. Sippell & H. R. Wong. 1971. The eastern pine-shoot borer, Eucosma gloriola (Lepidoptera: Tortricidae) in North America. Canadian Entomologist. 103: 1473-1486.
; Miller 1987a

Miller 1987a


Miller, W. E. 1987a. Guide to the Olethreutine moths of midland North America (Tortricidae). USDA Forest Service Agriculture Handbook 660: 1-104.
Eucopina spp.
Pinaceae Pinus banksiana Jack pine Eucopina gloriola Wong & Campbell 1967 ; Prentice 1966

Prentice 1966


Prentice, R. M. 1966. Vol. 4. Microlepidoptera. In: Forest Lepidoptera of Canada recorded by the Forest Insect Survey. Dept. For. Canada Publ. 1142: 543-840.
; De Groot et al. 1998

De Groot et al. 1998


De Groot, P., Grant, G.G., Nott, R.W., Langevin, D. 1998. Seasonal and diurnal flight activity of male Eucosma gloriola (Lepidoptera: Tortricidae). Canadian Entomologist. 130: 377-381.
Eucopina spp.
Pinaceae Pinus mugo dwarf mountain pine Eucopina gloriola Prentice 1966

Prentice 1966


Prentice, R. M. 1966. Vol. 4. Microlepidoptera. In: Forest Lepidoptera of Canada recorded by the Forest Insect Survey. Dept. For. Canada Publ. 1142: 543-840.
Eucopina spp.
Pinaceae Pinus nigra Austrian pine Eucopina gloriola Newman 1968

Newman 1968


Newman, J.H. 1968. First records of the white pine shoot borer, Eucosma gloriola (Lepidoptera: Olethreutidae) in Michigan. Mich. Entomol. 1: 267- 270.
Eucopina spp.
Pinaceae Pinus resinosa red pine Eucopina gloriola Prentice 1966

Prentice 1966


Prentice, R. M. 1966. Vol. 4. Microlepidoptera. In: Forest Lepidoptera of Canada recorded by the Forest Insect Survey. Dept. For. Canada Publ. 1142: 543-840.
Eucopina spp.
Pinaceae Pinus sp. pine Eucopina gloriola DeBoo et al. 1971

DeBoo et al. 1971


DeBoo, R. F., W. L. Sippell & H. R. Wong. 1971. The eastern pine-shoot borer, Eucosma gloriola (Lepidoptera: Tortricidae) in North America. Canadian Entomologist. 103: 1473-1486.
Eucopina spp.
Pinaceae Pinus strobus eastern white pine Eucopina gloriola Prentice 1966

Prentice 1966


Prentice, R. M. 1966. Vol. 4. Microlepidoptera. In: Forest Lepidoptera of Canada recorded by the Forest Insect Survey. Dept. For. Canada Publ. 1142: 543-840.
Eucopina spp.
Pinaceae Pinus sylvestris Scots pine Eucopina gloriola MacKay 1959

MacKay 1959


MacKay, M. R. 1959. Larvae of the North American Olethreutidae (Lepidoptera). Canadian Entomologist, Supplement 10: 1-338.
; Prentice 1966

Prentice 1966


Prentice, R. M. 1966. Vol. 4. Microlepidoptera. In: Forest Lepidoptera of Canada recorded by the Forest Insect Survey. Dept. For. Canada Publ. 1142: 543-840.
; Newman 1968

Newman 1968


Newman, J.H. 1968. First records of the white pine shoot borer, Eucosma gloriola (Lepidoptera: Olethreutidae) in Michigan. Mich. Entomol. 1: 267- 270.
; McKeague & Simmons 1978
Eucopina spp.
Pinaceae Pseudostuga sp. Douglas fir Eucopina gloriola DeBoo et al. 1971

DeBoo et al. 1971


DeBoo, R. F., W. L. Sippell & H. R. Wong. 1971. The eastern pine-shoot borer, Eucosma gloriola (Lepidoptera: Tortricidae) in North America. Canadian Entomologist. 103: 1473-1486.
; Miller 1987a

Miller 1987a


Miller, W. E. 1987a. Guide to the Olethreutine moths of midland North America (Tortricidae). USDA Forest Service Agriculture Handbook 660: 1-104.
Eucopina spp.
Pinaceae Pinus resinosa red pine Eucopina monitorana Lyons 1957

Lyons 1957


Lyons, L. A. 1957. Insects affecting seed production in red pine Ill. Eucosma moniforana Heinrich, Laspeyresia toreuta Grote (Lepidoptera: Olethreutidae), Rubsamenia sp. (Diptera: Cecidomyiidae), and other insects. Canadian Entomologist. 139: 150-164.
; Powell 1968

Powell 1968


Powell, J. A. 1968. Host associations and taxonomy of nearctic conifer cone moths in the genus Eucosma (Lepidoptera: Tortricidae). Hilgardia. 39: 1-36.
; Barras & Norris 1969

Barras & Norris 1969


Barras, S. J. & D. M. Norris. 1969. Bionomics of Eucosma monitorana (Lepidoptera: Tortricidae) attacking red pine cones in Wisconsin. Annals of the Entomological Society of America. 62: 1284-1290.
; Miller 1987a

Miller 1987a


Miller, W. E. 1987a. Guide to the Olethreutine moths of midland North America (Tortricidae). USDA Forest Service Agriculture Handbook 660: 1-104.
Eucopina spp.
Pinaceae Pinus sp. pine Eucopina monitorana Heinrich 1920, 1923b

Heinrich 1920, 1923b



Heinrich, C. 1920. On some forest Lepidoptera with descriptions of new species, larvae and pupae. Proceedings of the U.S. National Museum. 57: 53- 96.


Heinrich, C. 1923b. Revision of the North American moths of the subfamily Eucosminae of the family Olethreutidae. Bulletin of the United States National Museum. 123: 1-298.


Eucopina spp.
Pinaceae Pinus virginiana Virginia pine Eucopina monitorana Powell 1968

Powell 1968


Powell, J. A. 1968. Host associations and taxonomy of nearctic conifer cone moths in the genus Eucosma (Lepidoptera: Tortricidae). Hilgardia. 39: 1-36.
; Barras & Norris 1969

Barras & Norris 1969


Barras, S. J. & D. M. Norris. 1969. Bionomics of Eucosma monitorana (Lepidoptera: Tortricidae) attacking red pine cones in Wisconsin. Annals of the Entomological Society of America. 62: 1284-1290.
; Miller 1987a

Miller 1987a


Miller, W. E. 1987a. Guide to the Olethreutine moths of midland North America (Tortricidae). USDA Forest Service Agriculture Handbook 660: 1-104.
Eucopina spp.
Pinaceae Pinus jefferyi Jeffrey pine Eucopina ponderosa Powell 1968

Powell 1968


Powell, J. A. 1968. Host associations and taxonomy of nearctic conifer cone moths in the genus Eucosma (Lepidoptera: Tortricidae). Hilgardia. 39: 1-36.
; Cibrian-Tovar et al. 1986

Cibrian-Tovar et al. 1986


Cibrian-Tovar, D., B. H. Ebel, H. O. Yates & J. T. Mendez-Montiel (eds.). 1986. Cone and seed insects of the Mexican conifers. Southeastern Forest Experiment Station, U.S. Department of Agriculture.
Eucopina spp.
Pinaceae Pinus ponderosa Ponderosa pine Eucopina ponderosa Powell 1968

Powell 1968


Powell, J. A. 1968. Host associations and taxonomy of nearctic conifer cone moths in the genus Eucosma (Lepidoptera: Tortricidae). Hilgardia. 39: 1-36.
Eucopina spp.
Pinaceae Pinus albicaulis whitebark pine Eucopina rescissoriana Powell 1968

Powell 1968


Powell, J. A. 1968. Host associations and taxonomy of nearctic conifer cone moths in the genus Eucosma (Lepidoptera: Tortricidae). Hilgardia. 39: 1-36.
Eucopina spp.
Pinaceae Pinus attenuata knobcone pine Eucopina rescissoriana Powell 1968

Powell 1968


Powell, J. A. 1968. Host associations and taxonomy of nearctic conifer cone moths in the genus Eucosma (Lepidoptera: Tortricidae). Hilgardia. 39: 1-36.
Eucopina spp.
Pinaceae Pinus contorta lodgepole pine Eucopina rescissoriana Heinrich 1923b

Heinrich 1923b


Heinrich, C. 1923b. Revision of the North American moths of the subfamily Eucosminae of the family Olethreutidae. Bulletin of the United States National Museum. 123: 1-298.
; Powell 1968

Powell 1968


Powell, J. A. 1968. Host associations and taxonomy of nearctic conifer cone moths in the genus Eucosma (Lepidoptera: Tortricidae). Hilgardia. 39: 1-36.
Eucopina spp.
Pinaceae Pinus contorta var. murrayana Sierra lodgepole pine Eucopina rescissoriana Heinrich 1923b

Heinrich 1923b


Heinrich, C. 1923b. Revision of the North American moths of the subfamily Eucosminae of the family Olethreutidae. Bulletin of the United States National Museum. 123: 1-298.
; Heinrich 1920

Heinrich 1920


Heinrich, C. 1920. On some forest Lepidoptera with descriptions of new species, larvae and pupae. Proceedings of the U.S. National Museum. 57: 53- 96.
Eucopina spp.
Pinaceae Pinus monticola western white pine Eucopina rescissoriana Shea 1986

Shea 1986


Shea, P. J. 1986. Impact of insects on cone/seed production in three blister rust-resistant western white pine seed orchards. U.S. Department of Agriculture, Forest Service General Technical Report INT 203: 256-259.
; Powell 1968

Powell 1968


Powell, J. A. 1968. Host associations and taxonomy of nearctic conifer cone moths in the genus Eucosma (Lepidoptera: Tortricidae). Hilgardia. 39: 1-36.
Eucopina spp.
Pinaceae Abies concolor white fir Eucopina siskiyouana Heinrich 1923b

Heinrich 1923b


Heinrich, C. 1923b. Revision of the North American moths of the subfamily Eucosminae of the family Olethreutidae. Bulletin of the United States National Museum. 123: 1-298.
; Powell 1968

Powell 1968


Powell, J. A. 1968. Host associations and taxonomy of nearctic conifer cone moths in the genus Eucosma (Lepidoptera: Tortricidae). Hilgardia. 39: 1-36.
Eucopina spp.
Pinaceae Abies sp. fir Eucopina siskiyouana Heinrich 1923b

Heinrich 1923b


Heinrich, C. 1923b. Revision of the North American moths of the subfamily Eucosminae of the family Olethreutidae. Bulletin of the United States National Museum. 123: 1-298.
Eucopina spp.
Pinaceae Abies x shastensis Shasta red fir Eucopina siskiyouana Heinrich 1923b

Heinrich 1923b


Heinrich, C. 1923b. Revision of the North American moths of the subfamily Eucosminae of the family Olethreutidae. Bulletin of the United States National Museum. 123: 1-298.
Eucopina spp.
Pinaceae Picea engelmannii Engelmann spruce Eucopina sonomana Heinrich 1923b

Heinrich 1923b


Heinrich, C. 1923b. Revision of the North American moths of the subfamily Eucosminae of the family Olethreutidae. Bulletin of the United States National Museum. 123: 1-298.
Eucopina spp.
Pinaceae Pinus contorta lodgepole pine Eucopina sonomana Mitchell & Sower 1991

Mitchell & Sower 1991


Mitchell, R.G., Sower, L.L. 1991. Life history of the western pine shoot borer (Lepidoptera: Olethreutidae) and effects on lodgepole pine in central Oregon. Journal of Economic Entomology. 84: 206-211.
Eucopina spp.
Pinaceae Pinus ponderosa Ponderosa pine Eucopina sonomana Heinrich 1923b

Heinrich 1923b


Heinrich, C. 1923b. Revision of the North American moths of the subfamily Eucosminae of the family Olethreutidae. Bulletin of the United States National Museum. 123: 1-298.
; Thier & Marsden 1990

Thier & Marsden 1990


Thier, R W., Marsden, M.A. 1990. Vertical growth response of ponderosa pine infested by western pine shoot borer, Eucosma sonomana Kearfott (Lepidoptera: Olethreutidae). Canadian Entomologist 122: 343-347.
; Stevens & Jennings 1977

Stevens & Jennings 1977


Stevens, R.E., Jennings, D.T. 1977. Western pine-shoot borer: a threat to intensive management of Ponderosa pine in the Rocky Mountain area and southwest. U.S. Department of Agriculture, Forest Service General Technical Report (RM) 45: 1-8.
; Sower & Shorb 1984 ; Sower & Mitchell 1987

Sower & Mitchell 1987


Sower, L.L., Mitchell, R.G. 1987. Host-tree selection by western pine shoot borer (Lepidoptera: Olethreutidae) in ponderosa pine plantations. Environmental Entomology. 16: 1145-1147.
; Mitchell & Sower 1988, 1991

Mitchell & Sower 1988, 1991



Mitchell, R.G., Sower, L.L. 1988. Phenology and heat accumulation as field guides for the seasonal history of the western pine shoot borer (Lepidoptera: Olethreutidae). Journal of Economic Entomology. 81: 286-289.


Mitchell, R.G., Sower, L.L. 1991. Life history of the western pine shoot borer (Lepidoptera: Olethreutidae) and effects on lodgepole pine in central Oregon. Journal of Economic Entomology. 84: 206-211.


; Prueitt & Ross 1998

Prueitt & Ross 1998


Prueitt, S.C., Ross, D.W. 1998. Effects of environment and host genetics on Eucosma sonomana (Lepidoptera: Tortricidae) infestation levels. Environmental Entomology. 27: 1469-1472.
Eucopina spp.
Pinaceae Pinus sp. pine Eucopina sonomana Sartwell et al. 1980

Sartwell et al. 1980


Sartwell, C., G. E. Daterman, L. L. Sower, D. L. Overhulser & T. W. Koerber. 1980. Mating disruption with synthetic sex attractants controls damage by Eucosma sonomana (Lepidoptera: Tortricidae, Olethreutinae) in Pinus ponderosa plantations. 1. Manually applied polyvinyl chloride formulation. Canadian Entomologist. 112: 159-162.
; Sower et al. 1984, 1989

Sower et al. 1984, 1989



Sower, L. L., G. E. Daterman & C. Starwell. 1984. Surveying populations of western pine shoot borers (Lepidoptera: Olethreutidae). Journal of Economic Entomology. 77: 715-719.


Sower, L. L., G. E. Daterman & C. Starwell. 1989. Biology and impact of western pine shoot borer, Eucosma sonomana, in the western United States. In: Alfaro, R. I. & S. G. Glover (ed.), Insects affecting restoration: biology and damage. Forestry Canada, Pacific & Yukon Region, Victoria, Canada. 256 pp.


Eucopina spp.
Pinaceae Abies balsamea balsam fir Eucopina tocullionana Prentice 1966

Prentice 1966


Prentice, R. M. 1966. Vol. 4. Microlepidoptera. In: Forest Lepidoptera of Canada recorded by the Forest Insect Survey. Dept. For. Canada Publ. 1142: 543-840.
; Powell 1968

Powell 1968


Powell, J. A. 1968. Host associations and taxonomy of nearctic conifer cone moths in the genus Eucosma (Lepidoptera: Tortricidae). Hilgardia. 39: 1-36.
Eucopina spp.
Pinaceae Abies sp. fir Eucopina tocullionana Miller 1987a

Miller 1987a


Miller, W. E. 1987a. Guide to the Olethreutine moths of midland North America (Tortricidae). USDA Forest Service Agriculture Handbook 660: 1-104.
Eucopina spp.
Pinaceae Picea sp. spruce Eucopina tocullionana Heinrich 1920, 1923b

Heinrich 1920, 1923b



Heinrich, C. 1920. On some forest Lepidoptera with descriptions of new species, larvae and pupae. Proceedings of the U.S. National Museum. 57: 53- 96.


Heinrich, C. 1923b. Revision of the North American moths of the subfamily Eucosminae of the family Olethreutidae. Bulletin of the United States National Museum. 123: 1-298.


; MacKay 1959

MacKay 1959


MacKay, M. R. 1959. Larvae of the North American Olethreutidae (Lepidoptera). Canadian Entomologist, Supplement 10: 1-338.
; Powell 1968

Powell 1968


Powell, J. A. 1968. Host associations and taxonomy of nearctic conifer cone moths in the genus Eucosma (Lepidoptera: Tortricidae). Hilgardia. 39: 1-36.
; Miller 1987a

Miller 1987a


Miller, W. E. 1987a. Guide to the Olethreutine moths of midland North America (Tortricidae). USDA Forest Service Agriculture Handbook 660: 1-104.
Eucopina spp.
Pinaceae Pinus sp. pine Eucopina tocullionana Powell 1968

Powell 1968


Powell, J. A. 1968. Host associations and taxonomy of nearctic conifer cone moths in the genus Eucosma (Lepidoptera: Tortricidae). Hilgardia. 39: 1-36.
; Miller 1987a

Miller 1987a


Miller, W. E. 1987a. Guide to the Olethreutine moths of midland North America (Tortricidae). USDA Forest Service Agriculture Handbook 660: 1-104.
Eucopina spp.
Pinaceae Pinus strobus eastern white pine Eucopina tocullionana Prentice 1966

Prentice 1966


Prentice, R. M. 1966. Vol. 4. Microlepidoptera. In: Forest Lepidoptera of Canada recorded by the Forest Insect Survey. Dept. For. Canada Publ. 1142: 543-840.
; Powell 1968

Powell 1968


Powell, J. A. 1968. Host associations and taxonomy of nearctic conifer cone moths in the genus Eucosma (Lepidoptera: Tortricidae). Hilgardia. 39: 1-36.
; Ebel et al. 1975

Ebel et al. 1975


Ebel, B.H., Flavell, T.H., Drake, L.E., Yates, H.O., DeBarr, G.L. 1975. Seed and cone insects of southern pines. General technical Report/SE-8, USDA Forest Service, Southern Forest Experiment Station. 39 pp.
; De Groot 1998

De Groot 1998


De Groot, P. 1998. Life history and habits of the white pine cone borer, Eucosma tocullionana (Lepidoptera: Tortricidae). Canadian Entomologist. 130: 79-90.
; Powell 2006

Powell 2006


Powell, J. A. 2006. Database of Lepidoptera rearing lots, 1960-2005. University of California Berkeley, CA.
Eucopina spp.
Pinaceae Tsuga canadensis eastern hemlock Eucopina tocullionana Powell 1968

Powell 1968


Powell, J. A. 1968. Host associations and taxonomy of nearctic conifer cone moths in the genus Eucosma (Lepidoptera: Tortricidae). Hilgardia. 39: 1-36.
Eucopina spp.
Pinaceae Tsuga sp. hemlock Eucopina tocullionana USNM collection

USNM collection


USNM collection. Based on identified reared specimens in the collection of the National Museum of Natural History, Washington, D.C.
Eucopina spp.
PLANT FAMILY PLANT SPECIES PLANT COMMON NAME ASSOCIATED PEST REFERENCE LINK