The mesopleuron is the lateral part of the mesothorax and in Diptera comprises the majority of the lateral part of the thorax. It includes the sclerites between the anterior and posterior spiracles, which are the anepisternum, katepisternum, anepimeron, and katepimeron. This character refers to the predominant color of the mesopleuron, exclusive of the normal whitish areas (i.e., the dorsal and posterior margins of the anepisternum, the dorsal margin or dorsomedial spot on the katepisternum, and most of the katatergite and anatergite).

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