Abreu RLS, Fonseca CRV, Guerrero JCH, Paula EVCM. 2001. Preferência de vôo de nove espécies da família Scolytidae (Insecta: Coleoptera) na Amazônia Central. Acta Amazonica 31: 61–68. http://dx.doi.org/10.1590/1809-43922001311068.

Alonso-Zarazaga MA, Lyal CHC. 2009. A catalogue of family and genus group names in Scolytinae and Platypodinae with nomenclatural remarks (Coleoptera: Curculionidae). Zootaxa 2258: 1–134. https://doi.org/10.11646/zootaxa.2258.1.1

Atkinson, TH. 2019. Bark and ambrosia beetles. http://www.barkbeetles.info

Atkinson TH, Rabaglia RJ, Bright DE. 1990. Newly detected exotic species of Xyleborus (Coleoptera: Scolytidae) with a revised key to species in eastern North America. The Canadian Entomologist 122: 92–104. https://doi.org/10.4039/Ent12293-1

Atkinson TH, Carrillo D, Duncan RE, Peña JE. 2013. Occurrence of Xyleborus bispinatus (Coleoptera: Curculionidae: Scolytinae) Eichhoff in southern Florida. Zootaxa 3669: 96–100. https://doi.org/10.11646/zootaxa.3669.1.10


Barnouin T, Soldati F,  Roques A, Faccoli M, Kirkendall LR, Mouttet R, Daubree J-B, Noblecourt T (2020) Bark beetles and pinhole borers recently or newly introduced to France (Coleoptera: Curculionidae, Scolytinae and Platypodinae). Zootaxa: 4877: 51–74. https://doi.org/10.11646/zootaxa.4877.1.2

Bateman C, Kendra PE, Rabaglia R, Hulcr J. 2015. Fungal symbionts in three exotic ambrosia beetles, Xylosandrus amputatus, Xyleborinus andrewesi, and Dryoxylon onoharaense (Coleoptera: Curculionidae: Scolytinae: Xyleborini) in Florida. Symbiosis 66: 141–148. https://doi.org/10.1007/s13199-015-0353-z

Beaver RA. 1976. The biology of Samoan bark and ambrosia beetles (Coleoptera, Scolytidae and Platypodidae). Bulletin of Entomological Research 65: 531–548.

Beaver RA. 1987. The bark and ambrosia beetles (Coleoptera: Scolytidae and Platypodidae) of Tonga. New Zealand Entomologist 9: 64–70. https://doi.org/10.1080/00779962.1987.9722496

Beaver RA. 1988. Biological studies on ambrosia beetles of the Seychelles (Col, Scolytidae and Platypodidae). Journal of Applied Entomology 105: 62–73. https://doi.org/10.1111/j.1439-0418.1988.tb00162.x

Beaver RA. 1990. New records and new species of bark and ambrosia beetles from Thailand (Coleoptera: Scolytidae and Platypodidae). Deutsche Entomologische Zeitschrift 37: 279–284. https://doi.org/10.1002/mmnd.19900370408

Beaver RA. 1991. New synonymy and taxonomic changes in Pacific Scolytidae (Coleoptera). Annalen des Naturhistorisches Museums in Wien, 92B: 87–97.

Beaver RA. 1995a. New synonymy and taxonomic changes in Oriental and Australasian Scolytidae and Platypodidae (Insecta: Coleoptera). Annalen des Naturhistorischen Museums in Wien 97B: 197–204.

Beaver RA .1995b. Additions and corrections to the bark and ambrosia beetle fauna of Fiji (Coleoptera: Scolytidae). South Pacific Journal of Natural Science 14: 11‒26.

Beaver RA. 1998. New synonymy, new combinations, and taxonomic notes on Scolytidae and Platypodidae (Insecta: Coleoptera). Annalen des Naturhistorischen Museums in Wien, Series B, Botanik und Zoologie 100B: 179–192.

Beaver RA. 2005. A remarkable new species of Cyclorhipidion Hagedorn, and new records of bark and ambrosia beetles from Gabon (Coleoptera: Curculionidae, Scolytinae and Platypodinae). The Entomologist’s Monthly Magazine 141: 113–119.

Beaver RA. 2010. A review of the genus Hadrodemius Wood, with new synonymy and a key to species (Coleoptera: Curculionidae: Scolytinae). Zootaxa 2444: 51–57. https://doi.org/10.11646/zootaxa.2444.1.3

Beaver RA. 2011. New synonymy and taxonomic changes in bark and ambrosia beetles (Coleoptera: Curculionidae: Scolytinae, Platypodinae). Koleopterologische Rundschau 81: 277–289.

Beaver RA, Browne FG. 1975. The Scolytidae and Platypodidae (Coleoptera) of Thailand, a checklist with biological and zoogeographical notes. Oriental Insects 9: 283–311. https://doi.org/10.1080/00305316.1975.10434499

Beaver RA, Browne FG. 1978. The Scolytidae and Platypodidae (Coleoptera) of Penang, Malaysia. Oriental Insects 12: 575–624. https://doi.org/10.1080/00305316.1978.10432538

Beaver RA, Hulcr J. 2008. A review of the ambrosia beetle genus Cryptoxyleborus Schedl (Coleoptera, Curculionidae: Scolytinae). The Coleopterists Bulletin 62: 133–153. https://doi.org/10.1649/1026.1

Beaver RA, Liu L-Y. 2010. An annotated synopsis of Taiwanese bark and ambrosia beetles, with new synonymy, new combinations and new records (Coleoptera: Curculionidae: Scolytinae). Zootaxa 2602: 1–47. https://doi.org/10.11646/zootaxa.2602.1.1

Beaver RA, Liu L-Y. 2018. A synopsis of the bark and ambrosia beetles of Nepal with a key to the genera (Insecta: Coleoptera: Curculionidae: Platypodinae and Scolytinae). In: Hartmann M, Barclay MVL, Weipert J (Eds) Biodiversität und Naturausstattung im Himalaya VI. Verein der Freunde und Förderer des Naturkundemuseum Erfurt, Erfurt, pp. 521‒553.

Beaver RA, Löyttyniemi K. 1991. Annual flight patterns and diversity of bark and ambrosia beetles (Col., Scolytidae and Platypodidae) attracted to bait logs in Zambia. Journal of Applied Entomology 112: 505–511. https://doi.org/10.1111/j.1439-0418.1991.tb01084.x

Beaver RA, Smith SM. 2022. The bark and ambrosia beetles of Bhutan (Coleoptera: Curculionidae: Scolytinae and Platypodinae): a synopsis with three new species of Scolytinae. Zootaxa 5174: 1–24. https://doi.org/10.11646/zootaxa.5174.1.1

Beaver RA, Kajimura H, Goto H. 2008. Taxonomic changes and new records of Japanese bark and ambrosia beetles (Coleoptera, Curculionidae, Scolytinae). Elytra, Tokyo 36: 231–239.

Beaver RA, Sittichaya W, Liu L-Y. 2014. A synopsis of the scolytine ambrosia beetles of Thailand (Coleoptera: Curculionidae: Scolytinae). Zootaxa 3875: 1–82. http://dx.doi.org/10.11646/zootaxa.3875.1.1

Beaver RA, Sittichaya W, Liu L-Y. 2019. A review of the genus Immanus Hulcr & Cognato with a key to species (Coleoptera: Curculionidae: Scolytinae: Xyleborini). Zootaxa 4585: 378–386. http://dx.doi.org/10.11646/zootaxa.4585.2.11

Bedel LEM. 1888. Faune des Coléoptères du Bassin de la Seine. Vol. VI. Rhynchophora. Annales de la Société Entomologique de France (6) 7, Publication Hors Série, 385–444.

Beeson CFC. 1915. Notes on some Indian forest beetles. Indian Forester 16: 294–299.

Beeson CFC. 1929. Platypodidae and Scolytidae. In: Insects of Samoa, Part 4, Coleoptera, Fascicle 4: 217–248.

Beeson CFC. 1930. The biology of the genus Xyleborus, with more new species. Indian Forest Records 14: 209–272.

Beeson CFC. 1935a. Platypodidae and Scolytidae of the Society Islands. Bulletin of the Bernice P. Bishop Museum 142: 115–121.

Beeson CFC. 1935b. Scolytidae of the Marquesas. Bulletin of the Bernice P. Bishop Museum 142: 101–114.

Beeson CFC. 1940. Scolytidae and Platypodidae of the Mangarevan Expedition. Occasional Papers of the Bernice P. Bishop Museum 15: 191–203.

Beeson CFC. 1961. The ecology and control of the forest insects of India and the neighbouring countries. 2nd edition. Government of India, New Delhi, 767 pp.

Biedermann PHW. 2010. Observations on sex ratio and behavior of males in Xyleborinus saxesenii Ratzeburg (Scolytinae, Coleoptera). In: Cognato AI, Knížek M (Eds). Sixty years of discovering scolytine and platypodine diversity: A tribute to Stephen L. Wood. ZooKeys 56: 253–267. https://doi.org/10.3897/zookeys.56.530

Biedermann PHW, Taborsky M. 2011. Larval helpers and age polyethism in ambrosia beetles. Proceedings of the National Academy of Science USA 108: 17064–17069. https://doi.org/10.1073/pnas.1107758108

Biedermann PHW, Klepzig KR, Taborsky M. 2009. Fungus cultivation by ambrosia beetles: behavior and laboratory breeding success in three Xyleborine species. Environmental Entomology 38: 1096–1105. https://doi.org/10.1603/022.038.0417

Biedermann PHW, Klepzig KD, Taborsky M. 2011. Costs of delayed dispersal and alloparental care in the fungus-cultivating ambrosia beetle Xyleborus affinis Eichhoff (Scolytinae: Curculionidae). Behavioral Ecology Sociobiology 65: 1753–1761. https://doi.org/10.1007/s00265-011-1183-5

Blackburn T. 1885. [new taxa] In: Blackburn T, Sharp D Memoirs on the Coleoptera of the Hawaiian Islands. The Scientific Transactions of the Royal Dublin Society, Series 2, 3, 119–289, 300, pls. IV, V.

Blandford WFH. 1894a. Notes on Scolytidae and their food-plants. Insect Life 6: 260–265.

Blandford WFH. 1894b. The Rhynchophorous Coleoptera of Japan. Part III. Scolytidae. Transactions of the Entomological Society of London 1894: 53–141.

Blandford WFH. 1894c. Supplementary notes on the Scolytidae of Japan, with a list of species. Transactions of the Entomological Society of London 1894: 575–580. https://doi.org/10.1111/j.1365-2311.1894.tb02101.x

Blandford WFH. 1895. A list of the Scolytidae collected in Ceylon by Mr. George Lewis, with descriptions of new species. Annals and Magazine of Natural History, series 6, 15: 315–328. https://doi.org/10.1080/00222939508677888

Blandford WFH. 1896a. Contributions à la faune indo-chinoise 16e memoire. Annales de la Société Entomologique de France 65: 19–22.

Blandford WFH. 1896b. Descriptions of new Scolytidae from the Indo-Malayan and Austro-Malayan regions. Transactions of the Entomological Society of London 1896: 191–228. https://doi.org/10.1111/j.1365-2311.1896.tb00962.x

Blandford WFH. 1898. On some Oriental Scolytidae of economic importance, with descriptions of five new species. Transactions of the Entomological Society of London 1898: 423–430. https://doi.org/10.1111/j.1365-2311.1898.tb03298.x

Bohemann CH. 1859. Coleoptera. Species nova descripsit. Pp. 113–217, pls. I–II. In Virgin CH (ed): Konglika Svenska Fregatten Eugenies Resa omkring jorden under befall af C. A. Virgin, åren 1851–1853. Vetenskapliga iakttagelser på H. M. konung Oscar Den Förstes befallning utgifnaaf K. Svenska Vetenskaps Akademien. Andra Delen. Zoologi. 1. Insecta. Häft 4. Uppsala et Stockholm: P. A. Norstedt & Söner, Almquist et Wiksells, 614 pp., 9 pls.

Boieldieu M. 1859. Descriptions d’espèces nouvelles de coléoptères. Annales de la Société Entomologique de France, series 3, 7: 461–482, pl. 8.

Boland JM. 2016. The impact of an invasive ambrosia beetle on the riparian habitats of the Tijuana River Valley, California. Peer J 4: e2141. https://doi.org/:10.7717/peerj.2141

Bousquet Y. 2018. The dating of the fourth volume of Guillaume-Antoine Olivier's "Entomologie, ou histoire naturelle des insectes". ZooKeys 734: 137–148. https://doi.org/10.3897/zookeys.734.22901

Brader L. 1964. Étude de la relation entre le scolyte des rameaux du caféier, Xyleborus compactus Eichh (X morstatti Hag) et sa plante-hôte. Mededelingen van de Landbouwhogeschool Wageningen 64: 1–109.

Brar GS, Capinera JL, McLean S, Kendra PE, Ploetz RC, Peña JE. 2012. Effect of trap size, trap height, and age of lure on sampling Xyleborus glabratus (Coleoptera: Curculionidae: Scolytinae) and its flight periodicity and seasonality. Florida Entomologist 95: 1003–1011. https://doi.org/10.1653/024.095.0428

Brar GS, Capinera JL, Kendra PE, McLean S, Peña JE. 2013. Life cycle, development and culture of Xyleborus glabratus (Coleoptera: Curculionidae: Scolytinae). Florida Entomologist 96: 1158–1167. https://doi.org/10.1653/024.096.0357

Bright DE. 1968. Review of the tribe Xyleborini in America north of Mexico (Coleoptera: Scolytidae). The Canadian Entomologist 100: 1288–1323. https://doi.org/10.4039/Ent1001288-12

Bright DE. 1972. The Scolytidae and Platypodidae of Jamaica (Coleoptera). Bulletin of Institute of Jamaica, Science Series 21: 1–108.

Bright DE. 1985. Studies on West Indian Scolytidae (Coleoptera), 3: Checklist of Scolytidae of the West Indies with descriptions of new species and taxonomic notes. Entomologischen Arbeit aus dem Museum G. Frey 33–34: 169–188.

Bright DE, Rabaglia RJ. 1999. Dryoxylon, a new genus for Xyleborus onoharaensis Murayama, recently established in the southeastern United States (Coleoptera: Scolytidae). The Coleopterists Bulletin 53: 333–337.

Bright DE, Skidmore RE. 1997. A Catalog of Scolytidae and Platypodidae (Coleoptera), Supplement 1 (1990–1994). NRC Research Press, Ottawa, 368 pp.

Broun T. 1904. Descriptions of new genera and species of New Zealand Coleoptera. (Concluded). The Annals and Magazine of Natural History, series 7, 14: 105–127. https://doi.org/10.1080/03745480409442977

Browne FG. 1949. Notes on Malayan Scolytoidea (Coleoptera) with descriptions of new species. The Annals and Magazine of Natural History, series 12, 1: 892–912. https://doi.org/10.1080/00222934808653953

Browne FG. 1950. New Scolytidae and Platypodidae (Coleoptera) from Malaya. The Annals and Magazine of Natural History, series 12, 3: 641–650. https://doi.org/10.1080/00222935008654090

Browne FG. 1955. Synonymy and descriptions of some Oriental Scolytidae and Platypodidae (Coleoptera). Sarawak Museum Journal 6: 343–373.

Browne FG. 1958. Beetles: Scolytidae from Borneo. New species of Scolytomimus and Webbia (Scolytidae) from Borneo and Malaya. Sarawak Museum Journal 8: 487–497.

Browne FG. 1961a. Borer beetles from Bako National Park (Sarawak). Sarawak Museum Journal 10: 300–318.

Browne FG. 1961b. The biology of Malayan Scolytidae and Platypodidae. Malayan Forest Records 22: 1–255.

Browne FG. 1961c. The generic characters, habits, and taxonomic status of Premnobius Eichh. (Coleopt., Scolytidae). Report of the West African Timber Borer Research Unit, 4: 45–51.

Browne FG. 1962. Some Scolytidae and Platypodidae (Coleoptera) from the Oriental Region. Philippine Journal of Science 89: 201–220.

Browne FG. 1963. Taxonomic notes on Scolytidae (Coleoptera). Entomologische Berichten 23: 53–59.

Browne FG. 1965. On some Scolytidae and Platypodidae (Coleoptera), mainly from Africa and the Oriental region. Zoologische Mededelingen 40: 187–209.

Browne FG. 1966. Some Platypodidae and Scolytidae (Coleoptera) from the Philippine, Bismarck, and Solomon islands. Entomologiske Meddelelser 34: 233–257.

Browne FG. 1968a. Pests and diseases of forest plantation trees: An annotated list of the principal species occurring in the British Commonwealth. Clarendon Press, Oxford, xi + 1330 pp.

Browne FG. 1968b. A collection of Scolytidae and Platypodidae (Coleoptera) from Vietnam. The Entomologist’s Monthly Magazine 104: 133–134

Browne FG. 1972. Some oriental Scolytidae and Platypodidae (Coleoptera). Oriental Insects 6: 19–32. https://doi.org/10.1080/00305316.1972.10434049

Browne FG. 1974. A summary of the scolytid fauna (Coleoptera) of Fiji, with some new species. Commonwealth Forestry Review 53: 63–71.

Browne FG. 1980a. Bark and ambrosia beetles (Coleoptera, Scolytidae, and Platypodidae) intercepted at Japanese ports, with descriptions of new species I. Kontyû, 48: 370–379.

Browne FG. 1980b. Bark beetles and ambrosia beetles (Coleoptera, Scolytidae, and Platypodidae) intercepted at Japanese ports, with descriptions of new species, II. Kontyû 48: 380–389.

Browne FG. 1981. Bark and ambrosia beetles (Coleoptera, Scolytidae, and Platypodidae) intercepted at Japanese ports, with descriptions of new species V. Kontyû 49: 125–136.

Browne FG. 1983a. Bark beetles and ambrosia beetles (Coleoptera, Scolytidae, and Platypodidae) intercepted at Japanese ports, with descriptions of new species, VII. Kontyû 51: 554–572.

Browne FG. 1983b. Three new species of Scolytidae (Coleoptera) from southern China. The Entomologist's Monthly Magazine 119: 33–34.

Browne FG. 1984a. More new species of Scolytidae (Coleoptera) from Papua New Guinea. South Pacific Journal of Natural Science 6: 86–102.

Browne FG. 1984b. Some new species of Platypodidae and Scolytidae from Papua New Guinea. South Pacific Journal of Natural Science 4: 55–75.

Browne FG. 1986. Bark beetles and ambrosia beetles (Coleoptera, Scolytidae, and Platypodidae) intercepted at Japanese ports, with descriptions of new species, XIII. Kontyû 54: 89–99.

Bucini D, Balestra GM, Pucci C, Paparatti B. 2005. Bio-ethology of Anisandrus dispar F. and its possible involvement in dieback (Moria) diseases of hazelnut (Corylus avellana L.) plants in central Italy. Acta Horticulturae 686: 435–444. https://doi.org/10.17660/ActaHortic.2005.686.60

Bussler H, Immler T. 2007. Neue Borkenkäferarten in Bayern. Forstschutz Aktuell 38: 5–8.


Chararas C. 1962. Étude biologique des scolytides des conifères. Encyclopédie Entomologique, Ser A, 38. Lechevalier, Paris viii + 556 pp.

Chen Y, Dallara PL, Nelson LJ, Coleman TW, Hishinuma SM, Carrillo D, Seybold SJ. 2016. Comparative morphometric and chemical analyses of phenotypes of two invasive ambrosia beetles (Euwallacea spp.) in the United States of America. Insect Science 24: 647–662. https://doi.org/10.1111/1744-7917.12329

Choo HY. 1983. Taxonomic studies on the Platypodidae and Scolytidae (Coleoptera) from Korea. Doctoral Thesis. Seoul National University, Seoul, Korea, 128 pp.

Choo HY, Woo KS. 1985. A list of Korean bark and ambrosia beetles and their host plants. Korean Journal of Plant Protection 24: 163–167.

Cognato AI, Olson RO, Rabaglia RJ. 2011. An Asian ambrosia beetle, Xylosandrus amputatus (Blandford) (Curculionidae: Scolytinae: Xyleborini), discovered in Florida, U.S.A. The Coleopterists Bulletin 65: 43–45. https://doi.org/10.1649/0010-065X-65.1.43

Cognato AI, Rabaglia RJ, Vandenberg NJ. 2013. Another Asian ambrosia beetle, Xyleborinus artestriatus (Eichhoff 1878) (Coleoptera: Curculionidae: Scolytinae: Xyleborini), newly detected in North America. The Pan-Pacific Entomologist 89: 27–31. https://doi.org/10.3956/2012-53.1

Cognato AI, Hoebeke ER, Kajimura H, Smith SM. 2015. History of the exotic ambrosia beetles Euwallacea interjectus and Euwallacea validus (Coleoptera: Curculionidae: Xyleborini) in the United States. Journal of Economic Entomology 108: 1129–1135. https://doi.org/10.1093/jee/tov073

Cognato AI, Smith SM, Li Y, Pham T-H, Hulcr J. 2019. Genetic variability among native Xyleborus glabratus Eichhoff populations and the description of two related species. Journal of Economic Entomology 112: 1274–1284. https://doi.org/10.1093/jee/toz026

Cognato AI, Smith SM, Beaver RA. 2020a. Two new genera of Oriental xyleborine ambrosia beetles (Coleoptera, Curculionidae: Scolytinae). Zootaxa 4722: 540–554. https://doi.org/10.11646/zootaxa.4722.6.2

Cognato AI, Sari G, Smith SM, Beaver RA, Li Y, Hulcr J, Jordal BH, Kajimura H, Lin C-S, Pham TH, Singh S, Sittichaya W. 2020b. The essential role of taxonomic expertise in the creation of DNA databases for the identification and delimitation of Southeast Asian ambrosia beetles species (Coleoptera: Curculionidae: Scolytinae: Xyleborini). Frontiers in Ecology and Evolution 8: 27.https://doi.org/10.3389/fevo.2020.00027

Coleman TW, Poloni AL, Chen Y, Thu PQ, Li Q, Sun J, Rabaglia RJ, Man G, Seybold SJ. 2019. Hardwood injury and mortality associated with two shot hole borers, Euwallacea spp., in the invaded region of southern California, USA, and the native region of Southeast Asia. Annals of Forest Science 76: 61. https://doi.org/10.1007/s13595-019-0847-6

Colombari F, Martinez-Sañudo I, Battisti A. 2022. First report of the alien ambrosia beetle Cnestus mutilatus and further finding of Anisandrus maiche in the European part of the EPPO region (Coleoptera: Curculionidae: Scolytinae: Xyleborini) EPPO Bulletin, 00, 1–5. Available from: https://doi.org/10.1111/epp.12840

Cooperband MF, Stouthamer R, Carillo D, Eskalen A, Thibault T, Cossé AA, Castrillo LA, Vandenberg JD, Rugman-Jones PF. 2016. Biology of two members of the Euwallacea fornicatus species complex (Coleoptera: Curculionidae: Scolytinae), recently invasive in the U.S.A., reared on an ambrosia beetle artificial diet. Agricultural and Forest Entomology 18: 223–237. https://doi.org/10.1111/afe.12155

Coyle DR, Booth DC, Wallace MS. 2005. Ambrosia beetle (Coleoptera: Scolytidae) species, flight, and attack on living eastern cottonwood trees. Journal of Economic Entomology 98: 2049–2057. https://doi.org/10.1603/0022-0493-98.6.2049


Davis MA, Dute RR. 1997. Fungal associates of the Asian ambrosia beetle, Xylosandrus crassiusculus. Southern Nursery Association Research Conference 42: 106–112.

Dodelin B. 2018. Espèce invasive nouveau pour la faune de France Scolyte Cyclorhipidion fukiense installé en Europe. https://entomodata.wordpress.com/2018/04/24/cyclorhipidion-fukiense-installe-en-europe/

Dodelin B (2019) Ambrosiophilus atratus est arrivé dans l’Est de la France. https://entomodata.wordpress.com/2019/09/13/ambrosiophilus-atratus-est-arrive-dans-lest-de-la-france/

Dole SA, Beaver RA. 2008. A review of the Australian species of Xylosandrus Reitter (Coleoptera: Curculionidae: Scolytinae). The Coleopterists Bulletin 62: 481–492. http://dx.doi.org/10.1649/1108.1

Dole SA, Cognato AI, 2010. Revision of Xylosandrus Reitter (Curculionidae: Scolytinae). Proceedings of the California of Science 61: 451–545.

Drake CJ. 1921. A new ambrosia beetle from the Adirondacks: notes on the work of Xyloterinus politus Say. Ohio Journal of Science 21: 201–205.

Dugès E. 1888. Métamorphoses de quelques Coléoptères du Mexique. Annales de la Société entomologique de Belgique 31: 137–147, 2 pls.


Egger A. 1973. Beiträge zur Biologie und Bekämpfung von Xyleborus (Anisandrus) dispar F. und X. saxeseni Ratz (Col, Scolytidae). Anzeiger für Schädlingskunde Pflanzen- und Umweltschutz 46: 183–186. https://doi.org/10.1007/BF01991812

Eggers H. 1920. 60 neue Borkenkäfer (Ipidae) aus Afrika, nebst zehn neuen Gattungen, zwei Abarten. (Schluss). Entomologische Blätter 16: 33–45.

Eggers H. 1922. Seltene und neue paläarktische Borkenkäfer, III. Entomologische Blätter 18: 12–18.

Eggers H. 1923. Neue indomalayische Borkenkäfer (Ipidae). Zoologische Mededeelingen 7: 129–220.

Eggers H. 1925. Ipidae aus Birma. Sborník Entomologického Oddělení Národního Musea v Praze 3: 151–160.

Eggers H. 1926. Japanische Borkenkäfer, I. Entomologische Blätter 22: 145–148.

Eggers H. 1927a. Neue indomalayische Borkenkäfer (Ipidae). I. Nachtrag. Treubia 9: 390–408.

Eggers H. 1927b. Neue indomalayische Borkenkäfer (Ipidae), II. Nachtrag. Philippine Journal of Science 33: 67–108.

Eggers H. 1929. Zur Synonymie der Borkenkäfer (Ipidae, Col) I. Wiener Entomologische Zeitung 46: 41–55.

Eggers H. 1930. Neue Xyleborus-Arten (Col. Scolytidae) aus Indien. Indian Forest Records, Entomology Series 14: 177–208.

Eggers H. 1932. Neue Borkenkafer (Ipidae, Col.) aus Africa (Nachtrag V.). Revue de Zoologie et de Botanique Africaines 22:191–304.

Eggers H. 1933a. Borkenkäfer (Ipidae, Col) aus Südamerika, VI. Material des Muséum Paris aus Franz. Guayana und Venezuela. Travaux du Laboratoire d'Entomologie, Muséum National d'Histoire Naturelle, Mémoires Originaux 1: 1–37.

Eggers H. 1933b. Zur paläarktischen Borkenkäferfauna, I. Entomologische Blätter 29: 49–56.

Eggers H. 1934a. Borkenkäfer (Ipidae, Col) aus Sudamerika, VII. Entomologische Blätter 30: 78–84.

Eggers H. 1934b. Zur Synonymie der Borkenkäfer (Ipidae, Col) IV. Entomologisches Nachrichtenblatt 8: 25–29.

Eggers H, 1935. Borkenkäfer aus Südamerika (Ipidae, Col) (Fortsetzung). VII. Vergessene und neue Gattungen (1. Teil). Revista de Entomologia, Rio de Janeiro 5: 153–159.

Eggers H. 1936a. Neue Borkenkäfer (Scolytidae, Col) aus Indien. The Annals and Magazine of Natural History, series 10, 17: 626–636. https://doi.org/10.1080/00222933608655163

Eggers H. 1936b. Neue indomalayische Borkenkäfer (Ipidae) III Nachtrag. Tijdschrift voor Entomologie 79: 77–91.

Eggers H. 1937. Zur paläarktischen Borkenkäferfauna, IV. Entomologische Blätter 33: 334–335.

Eggers H. 1939a. Entomological results from the Swedish expedition 1934 to Burma and British India. Coleoptera: Ipidae, gesammelt von René Malaise. Arkiv för Zoologi 31A: 1–14.

Eggers H. 1939b. Japanische Borkenkäfer, II. Arbeiten über Morphologische und Taxonomische Entomologie aus Berlin-Dahlem 6: 114–123.

Eggers H. 1940. Neue indomalayische Borkenkäfer (Ipidae) III Nachtrag (Forstsetzung). Tijdschrift voor Entomologie 83: 132–154.

Eggers H. 1941a. Borkenkäfer aus Südamerika. (Coleoptera: Ipidae). IX. Insel Guadeloupe. Arbeiten über Morphologische und Taxonomische Entomologie aus Berlin-Dahlem 8: 99–109.

Eggers H. 1941b. Neue Borkenkäfer (Ipidae, Col) aus China. Entomologische Blätter 37: 222–226.

Eggers H. 1942. Zur palaearktischen Borkenkäferfauna (Coleoptera: Ipidae). VIII. Borkenkäfer aus dem asiatischen Russland. Arbeiten über Morphologische und Taxonomische Entomologie aus Berlin-Dahlem 9: 27–36.

Eggers H. 1943. Ipidae (Scolytidae) (Coleoptera Phytophaga). In: Exploration du Parc National Albert, I. Mission C. F. De Witte 1933–1935, Fascicle 43: 63–68.

Eggers H. 1944. Zur paläarktischen Borkenkäferfauna (Coleoptera, Ipidae) X. Entomologische Blätter 40: 140–143.

Eichhoff WJ, 1864. Uber die Mundtheile und die Fühlerbildung der europäischen Xylophagi sens strict. Berliner Entomologische Zeitschrift 8: 17–46. https://doi.org/10.1002/mmnd.18640080103

Eichhoff WJ. 1866. Ueber einige Bostrichiden. Berliner Entomologische Zeitschrift 10: 275–278. https://doi.org/10.1002/mmnd.18660100118

Eichhoff WJ. 1868a. Neue amerikanische Borkenkäfer-Gattungen und Arten. Berliner Entomologische Zeitschrift 11: 399–402. https://doi.org/10.1002/mmnd.18670110320

Eichhoff WJ. 1868b. Neue amerikanische Borkenkäfer-Gattung und Arten. Berliner Entomologische Zeitschrift 12: 145–152. https://doi.org/10.1002/mmnd.18680120213

Eichhoff WJ. 1869. Neue exotische Xyleborus-Arten. Berliner Entomologische Zeitschrift 12: 273–280. http://dx.doi.org/10.1002/mmnd.18680120214

Eichhoff WJ 1876a. Tomicides. In: Chapuis F & Eichhoff WJ, Scolytides recueillis au Japon par M. G. Lewis. Annales de la Société Entomologique de Belgique 18: 195–204.

Eichhoff WJ. 1876b. Synonymisches über Tomiciden. Stettiner Entomogische Zeitung 37: 378–379.

Eichhoff WJ. 1877. Japanische Scolytidae. Deutsche Entomologische Zeitschrift 21: 117–128.

Eichhoff WJ. 1878a. Neue oder noch unbeschriebene Tomicinen. Stettiner Entomologische Zeitung 39: 383–392.

Eichhoff WJ. 1878b.  Ratio, descriptio, emendatio eorum Tomicinorum qui sunt in Dr medic. Chapuisii et autoris ipsius collectionibus et quos praeterea recognovit scriptor. Mémoires de la Société Royale des Sciences de Liège, Série 2e, 8: 1–531 + pl. I–V.

Eichhoff WJ. 1880. Description of a new species of the family Scolytidae from Sumatra. Notes from the Leyden Museum 2: 189–190.

Eichhoff WJ. 1886. Zwei neue ost-indische Scolytiden-Gattungen. Notes from the Leyden Museum 8: 24–26.

Entwhistle PF. 1972. Pests of cocoa. Longman, London, 779 pp.

Erichson WF. 1836. Systematische Auseinandersetzung der Familie der Borkenkäfer (Bostrichidae). Archiv für Naturgeschichte 2: 45–65.

Erichson WF. 1842. Beitrag zur Insecten-Fauna von Vandiemensland, mit besonderer Berücksichtigung der geographischen Verbreitung der Insecten. Archiv für Naturgeschichte 8: 83–287, 2 pls. https://doi.org/10.5962/bhl.part.21656

Eskalen A, Stouthamer R, Lynch SC, Rugman-Jones PF, Twizeyimana M, Gonzalez A, Thibault T. 2013. Host range of Fusarium dieback and its ambrosia beetle (Coleoptera: Scolytinae) vector in southern California. Plant Disease 97: 938–951. https://doi.org/10.1094/PDIS-11-12-1026-RE


Fabricius JC. 1775. Systema Entomologiae, sistens insectorum classes, ordines, genera, species adiectis synonymis, locis, descriptionibus, observationibus. Flensburgi et Lipsiae: Officina Libraria Kortii, xxxii + 832 pp. https://doi.org/10.5962/bhl.title.36510

Fabricius JC. 1792. Entomologia systematica emendata et aucta. Secundum classes, ordines, genera, species adjectis synonimis, locis, observationibus, descriptionibus. Tomus I. Pars II. Hafniae: C.G. Proft, xx + 538 pp. https://doi.org/10.5962/bhl.title.122153

Fabricius JC. 1801. Systema eleutheratorum secundum ordines, genera, species; adiectis synonimis, locis, observationibus, descriptionibus, Tomus II. Kiliae: Bibliopoli Academici Novi, 687 pp.

Faccoli M. 2008. First record of Xyleborus atratus Eichhoff from Europe with an illustrated key to the European Xyleborini (Coleoptera: Curculionidae: Scolytinae). Zootaxa 1772: 55–62. https://doi.org/10.11646/zootaxa.1772.1.2

Faccoli M, Frigimelica G, Mori N, Toffolo EP, Vettorazzo M, Simonato M. 2009. First record of Ambrosiodmus (Hopkins, 1915) (Coleoptera, Curculionidae, Scolytinae) in Europe. Zootaxa 2303: 57–60. https://doi.org/10.11646/zootaxa.2303.1.4

Ferrari JA. 1867. Die Forst- und Baumzuchtschädlichen Borkenkäfer (Tomicides Lac.) aus der Familie der Holzverderber (Scolytides Lac.), mit besonderer Berücksichtigung vorzüglich der europäischen Formen, und der Sammlung des k. k. zoologischen Kabinets in Wien. Wien: Carl Gerold's Sohn, Wien 2 + 96 pp.

Fischer M. 1954. Untersuchungen über den Kleinen Holzbohrer (Xyleborinus saxsseni Ratz). Plflanzenschutzberichte 12: 137–180.

Flechtmann CAH, Cognato AI. 2011. First report of Amasa truncata (Coleoptera: Curculionidae: Scolytinae) in Brazil. The Coleopterists Bulletin 65: 417–421. https://doi.org/10.1649/072.065.0419

Flechtmann CAH, Atkinson TH. 2016. First records of Xylosandrus crassiusculus (Motschulsky) (Coleoptera: Curculionidae: Scolytinae) from South America, with notes on its distribution and spread in the New World. The Coleopterists Bulletin 70: 79–83. https://doi.org/10.1649/072.070.0109

Formby JP, Krishnan N, Riggins JJ. 2013. Supercooling in the redbaby ambrosia beetle (Coleoptera: Curculionidae). The Florida Entomologist 96: 1530–1540. http://dx.doi.org/10.1653/024.096.0435

Fraedrich SW, Harrington TC, Rabaglia RJ, Ulyshen MD, Mayfield III, AE, Hanula JL, Eickwort JM, Miller DR. 2008. A fungal symbiont of the redbay ambrosia beetle causes a lethal wilt in redbay and other Lauraceae in the Southeastern United States. Plant Disease 92: 215–224. https://doi.org/10.1094/PDIS-92-2-0215

Freeman S, Sharon M, Maymon M, Mendel Z, Protasov A, Aoki T, Eskalen A, O’Donnell K. 2013. Fusarium euwallaceae sp. nov.—a symbiotic fungus of Euwallacea sp., an invasive ambrosia beetle in Israel and California. Mycologia 105: 1595–1606. https://doi.org/10.3852/13-066

French JR, Roeper RA. 1975. Studies on the biology of the ambrosia beetle Xyleborus dispar (F) (Coleoptera: Scolytidae). Zeitschrift für Angewandte Entomologie 78: 241–247. https://doi.org/10.1111/j.1439-0418.1975.tb04178.x


Galko J, Nikolov C, Kimoto T, Kunca A, Gubka A, Vakula J, Zúbrik M, Ostriboň M. 2014. Attraction of ambrosia beetles to ethanol baited traps in a Slovakian oak forest. Biologia. 69: 1376–1383. https://doi.org/10.2478/s11756-014-0443-z

Gallego D, Lencina JL, Mas H, Ceveró J, Faccoli M. 2017. First record of the granulate ambrosia beetle, Xylosandrus crassiusculus (Coleoptera: Curculionidae: Scolytinae), in the Iberian Peninsula. Zootaxa 2743: 431–434. http://dx.doi.org/10.11646/zootaxa.4273.3.7

García-Avila CDJ, Trujillo-Arriaga FJ, López-Buenfil JA, González-Gómez R, Carrillo D, Cruz LF, Ruiz-Galván I, Quezada-Salinas A, Acevedo-Reyes N. 2016. First report of Euwallacea nr. fornicatus (Coleoptera: Curculionidae) in Mexico. Florida Entomologist 99: 555–556. https://doi.org/10.1653/024.099.0335

Garonna AP, Dole SA, Saracino A, Mazzoleni S, Cristinzio G. 2012. First record of the black twig borer Xylosandrus compactus (Eichhoff) (Coleoptera: Curculionidae, Scolytinae) from Europe. Zootaxa 3251: 64–68. https://doi.org/10.11646/zootaxa.3251.1.5

Gohli J, Selvarajah T, Kirkendall LR, Jordal BH. 2016. Globally distributed Xyleborus species reveal recurrent intercontinental dispersal in a landscape of ancient worldwide distributions. BMC Evolutionary Biology 16: 37. https://doi.org/10.1186/s12862-016-0610-7

Gohli J, Kirkendall LR, Smith SM, Cognato AI, Hulcr J, Jordal BH. 2017. Biological factors contributing to bark and ambrosia beetle species diversification. Evolution 71: 1258–1272. https://doi.org/10.1111/evo.13219

Gómez D, Suárez M, Martínez G. 2017. Amasa truncata (Erichson) (Coleoptera: Curculionidae: Scolytinae): A new exotic ambrosia beetle in Uruguay. The Coleopterists Bulletin 71: 825–826. https://doi.org/10.1649/0010-065X-71.4.825

Gomez DF, Rabaglia RJ, Fairbanks KEO, Hulcr J. 2018a. North American Xyleborini north of Mexico: a review and key to genera and species (Coleoptera, Curculionidae, Scolytinae). ZooKeys 768: 19–68. https://doi.org/10.3897/zookeys.768.24697

Gomez DF, Skelton J, Steininger MS, Stouthamer R, Rugman-Jones P,  Sittichaya W, Rabaglia RJ, Hulcr J. 2018b. Species within the Euwallacea fornicatus (Coleoptera: Curculionidae) complex revealed by morphometric and phylogenetic analyses. Insect Systematics and Diversity 2(6): 2, 1–11. https://doi.org/10.1093/isd/ixy018

Gray B, Wylie FR. 1974. Forest tree and timber insect pests in Papua New Guinea II. Pacific Insects 16: 67–115.


Haack R. 2006. Exotic bark- and wood-boring Coleoptera in the United States: recent establishments and interceptions. Canadian Journal of Forest Research 36: 269–288. https://doi.org/10.1139/x05-249

Hagedorn M. 1904. Enumeratio Scolytidarum e Sikkim et Japan natarum Musei historico-naturalis Parisiorum, quas dominus J. Harmand annis 1890 et 1901 collegit descriptionibus specierum novarum adjectis. Bulletin du Muséum d'Histoire Naturelle 10: 122–126.

Hagedorn M. 1908. Diagnosen bisher unbeschriebener Borkenkäfer. Erste Serie. Deutsche Entomologische Zeitschrift 1908: 369–382.

Hagedorn M. 1909. Diagnosen bisher unbeschriebener Borkenkäfer (Col). Deutsche Entomologische Zeitschrift 1909: 733–746.

Hagedon M. 1910a. Diagnosen bisher unbeschriebener Borkenkäfer (Col). Zweite Serie, zweite Hälfte. Deutsche Entomologische Zeitschrift 1910: 1–13.

Hagedorn M. 1910b. Ipidae. In: Schenkling, S. (Ed.). Coleopterorum Catalogus auspiciis et auxilio. Pars 4. W. Junk, Den Haag, pp. 1–134. https://doi.org/10.1007/978-94-011-9697-0_1

Hagedorn M. 1912a. Ipiden als Kaffeeschädlinge. Entomologische Blätter 8: 33–43.

Hagedorn M. 1912b. Neue Borkenkäfergattungen und Arten aus Afrika (Col.). Deutsche Entomologische Zeitschrift 1912: 351–356, pls. 6–7.

Halbert SE. 2011. Entomology Section. Tri-ology 50: 6–7.

Hanula JL, Mayfield III AE, Fraedrich SW, Rabaglia RJ. 2008. Biology and host associations of redbay ambrosia beetle (Coleoptera: Curculionidae: Scolytinae), exotic vector of laurel wilt killing redbay trees in southeastern United States. Journal of Economic Entomology 101: 1276–1286. https://doi.org/10.1603/0022-0493(2008)101[1276:BAHAOR]2.0.CO;2

Harrington TC, Yun HY, Lu SS, Goto H, Aghayeva DN, Fraedrich SW. 2011. Isolations from the redbay ambrosia beetle, Xyleborus glabratus, confirm that the laurel wilt pathogen Raffaelea lauricola, originated in Asia. Mycologia 103: 1028–1036. https://doi.org/10.3852/10-417

Hauptman T, Piškur B, Faccoli M, Rekanje B, Marin Č, Jurc M. 2019. The first record of two non-native ambrosia beetles in Slovenia: Ambrosiodmus rubricollis (Eichhoff, 1875) and Ambrosiophilus atratus (Eichhoff, 1875) (Coleoptera: Curculionidae, Scolytinae). Zootaxa 4657: 397–400. http://dx.doi.org/10.11646/zootaxa.4657.2.13

Hoebeke ER. 1991. An Asian ambrosia beetle, Ambrosiodmus lewisi, new to North America (Coleoptera: Scolytidae). Proceedings of the Entomological Society of Washington 93: 420–424. https://doi.org/10.4289/07-048.1

Hoebeke ER, Rabaglia RJ. 2007. First reported occurrence of Xyleborinus alni (Coleoptera: Curculionidae: Scolytinae) in the eastern United States, with notes on its recognition and tree hosts. Proceedings of the Entomological Society of Washington 109: 240–248.

Hoebeke ER, Rabaglia RJ. 2008. Xyleborus seriatus Blandford (Coleoptera: Curculionidae: Scolytinae), an Asian ambrosia beetle new to North America. Proceedings of the Entomological Society of Washington 110: 470–476.

Hoebeke ER, Rabaglia RJ, Knížek M, Weaver JS. 2018. First records of Cyclorhipidion fukiense (Eggers) (Coleoptera: Curculionidae: Scolytinae: Xyleborini), an ambrosia beetle native to Asia, in North America. Zootaxa 4394: 243–250. https://doi.org/10.11646/zootaxa.4394.2.7

Hoffman CE. 1941. Biological observations of Xylosandrus germanus (Bldfd.). Journal of Economic Entomology 34: 38–42. https://doi.org/10.1093/jee/34.1.38

Hopkins AD. 1915a. Classification of the Cryphalinae with descriptions of new genera and species. United States Department of Agriculture, Report No. 99. Washington: Government Printing Office, 75 pp., 4 pls. https://doi.org/10.5962/bhl.title.65905

Hopkins AD. 1915b. Contributions toward a monograph of the scolytid beetles, Part II. Preliminary classification of the superfamily Scolytoidea. United States Department of Agriculture, Technical Series, No. 17. Washington: Government Printing Office, vi + 165–232, pls. IX–XV.

Hosking GP. 1973. Xyleborus saxeseni, its life history and flight behavior in New Zealand. New Zealand Journal of Forestry Science 3: 37–53.

Hubbard HG. 1897. The ambrosia beetles of the United States. Bulletin of the United States Department of Agriculture, Division of Entomology, new series, 7: 9‒30.

Hulcr J. 2010. Taxonomic changes in palaeotropical Xyleborini (Coleoptera: Curculionidae: Scolytinae). ZooKeys 56: 105–119.  https://doi.org/10.3897/zookeys.56.520

Hulcr J, Cognato AI. 2009. Three new genera of Oriental Xyleborini (Coleoptera: Curculionidae: Scolytinae). Zootaxa 2204: 19–36. https://doi.org/10.11646/zootaxa.2204.1.2

Hulcr J, Cognato AI. 2010a. New genera of Palaeotropical Xyleborini (Coleoptera: Curculionidae: Scolytinae) based on congruence between morphological and molecular characters. Zootaxa 2717: 1–33. https://doi.org/10.11646/zootaxa.2717.1.1

Hulcr J, Cognato AI. 2010b. Repeated evolution of theft in fungus farming ambrosia beetles. Evolution 64: 3205–3212. https://doi.org/10.1111/j.1558-5646.2010.01055.x

Hulcr J, Cognato AI. 2013. Xyleborini of New Guinea: a taxonomic monograph. Thomas Say Publications in Entomology, Entomological Society of America, Lanham, Maryland, 176 pp.

Hulcr J, Lou Q-Z. 2013. The redbay ambrosia beetle (Coleoptera: Curculionidae) prefers Lauraceae in its native range: records form the Chinese National Insect Collection. Florida Entomologist 96: 1595–1597. https://doi.org/10.1653/024.096.0444

Hulcr J, Mann R, Stelinski LL. 2011. The scent of a partner: ambrosia beetles are attracted to volatiles from their fungal symbiont. Journal of Chemical Ecology 37: 1374–1377. https://doi.org/10.1007/s10886-011-0046-x

Hulcr J, Dole SA, Beaver RA, Cognato AI. 2007. Cladistic review of generic taxonomic characters in Xyleborini (Coleoptera: Curculionidae: Scolytinae). Systematic Entomology 32: 568–584. https://doi.org/10.1111/j.1365-3113.2007.00386.x

Hulcr J, Atkinson TH, Cognato AI, Jordal BH, McKenna DD. 2015. Morphology, taxonomy, and phylogenetics of bark beetles. In: Vega FE, Hofstetter RW (Eds) Bark beetles: biology and ecology of native and invasive species. Elsevier, San Diego, CA, 41−84.

Hulcr J, Black A, Prior K, Chen C-Y, Li H-F. 2017. Studies of ambrosia beetles in their native range help predict invasion impact. Florida Entomologist 100: 257–261. https://doi.org/10.1653/024.100.0219


Ito M, Kajimura H, Hamaguchi K, Araya K, Lakatos F. 2008. Genetic structure of Japanese populations of an ambrosia beetle, Xylosandrus germanus (Curculionidae: Scolytinae). Entomological Science 11: 375–383. https://doi.org/10.1111/j.1479-8298.2008.00280.x


Johnson AJ, McKenna DD, Jordal BH, Cognato AI, Smith SM, Lemmon AR, Moriarty Lemmon EL, Hulcr J. 2018. Phylogenomics clarifies repeated evolutionary origins of inbreeding and fungus farming in bark beetles (Curculionidae, Scolytinae). Molecular Phylogenetics and Evolution 127: 229–238. https://doi.org/10.1016/j.ympev.2018.05.028

Jordal BH. 2002. Elongation factor 1 a resolves the monophyly of the haplodiploid ambrosia beetles Xyleborini (Coleoptera: Curculionidae). Insect Molecular Biology 11: 453–465. https://doi.org/10.1046/j.1365-2583.2002.00354.x

Jordal BH, Cognato AI. 2012. Molecular phylogeny of bark and ambrosia beetles reveals multiple origins of fungus farming during periods of global warming. BMC Evolutionary Biology 12: 133. https://doi.org/10.1186/1471-2148-12-133

Jordal BH, Normark BB, Farrell BD. 2000. Evolutionary radiation of an inbreeding haplodiploid beetle lineage (Curculionidae, Scolytidae). Biological Journal of the Linnean Society 71: 483–499.  https://doi.org/10.1111/j.1095-8312.2000.tb01270.x

Jordal BH, Smith SM, Cognato AI. 2014. Classification of weevils as a data-
driven science: leaving opinion behind. ZooKeys 439: 1−18.



Kajimura H, Hijii N. 1992. Dynamics of the fungal symbionts in the gallery system and the mycangia of the ambrosia beetle, Xylosandrus mutilatus (Blandford) (Coleoptera: Scolytidae) in relation to its life history. Ecological Research 7: 107–117. https://doi.org/10.1007/BF02348489

Kajimura H, Hijii N. 1994. Reproduction and resource utilization of the ambrosia beetle Xylosandrus mutilatus, in field and experimental populations. Entomologia Experimentalis et Applicata 71: 121–132. https://doi.org/10.1111/j.1570-7458.1994.tb01778.x

Kalshoven LGE. 1958. Studies on the biology of Indonesian Scolytoidea I. Xyleborus fornicatus Eichh as a primary and secondary shot-hole borer in Java and Sumatra. Entomologische Berichten, Amsterdam 18: 147–160; 185–193.

Kalshoven LGE. 1959a. New cases of synonymy in Indomalayan scolytids. Entomologische Berichten, Amsterdam 19: 93–97.

Kalshoven LGE. 1959b. Studies on the biology of Indonesian Scolytoidea 4. Data on the habits of Scolytidae. Second part. Tijdschrift voor Entomologie 102: 135–173 + pls. 15–22.

Kalshoven LGE. 1960. Two new cases of synonymy in Indomalayan Platypodidae and Scolytidae. Entomologische Berichten, Amsterdam 20: 63–64.

Kalshoven LGE. 1961. A study of the twig borer Xyleborus morigerus Blandford, mainly based on observations in Java. Tijdschrift voor Entomologie 104: 93–110.

Kalshoven LGE. 1962. Note on the habits of Xyleborus destruens Bldf, the near-primary borer of teak trees on Java. Entomologische Berichten, Amsterdam 22: 7–18.

Kalshoven LGE. 1964. The occurrence of Xyleborus perforans (Woll) and X. similis in Java (Coleoptera, Scolytidae). Beaufortia 11: 131–142.

Kasson MT, Wickert KL, Stauder CM, Macias AM, Berger MC, Simmons DR, Short DPG, DeVallance DB, Hulcr J. 2016. Mutualism with aggressive wood-degrading Flavodon ambrosius (Polyporales) facilitates niche expansion and communal social structure in Ambrosiophilus ambrosia beetles. Fungal Ecology 23: 86–96. https://doi.org/10.1016/j.funeco.2016.07.002

Kendra PE, Montgomery WS, Niogret J, Deyrup MA, Guillen L, Epsky ND. 2012. Xyleborus glabratus, X. affinis, and X. ferrugineus (Coleoptera: Curculionidae: Scolytinae): Electroantennogram responses to host-based attractants and temporal patterns in host-seeking flight. Environmental Entomology 41: 1597–1605. https://doi.org/10.1603/EN12164

Kendra PE, Niogret J, Montgomery WS, Deyrup MA, Epsky ND. 2015. Cubeb oil lures: terpenoid emissions, trapping efficacy, and longevity for attraction of redbay ambrosia beetle (Coleoptera: Curculionidae: Scolytinae). Journal of Economic Entomology 108: 350–361. https://doi.org/10.1093/jee/tou023

Kendra PE, Montgomery WS, Deyrup MA, Wakarchuk D. 2016. Improved lure for redbay ambrosia beetle developed by enrichment of α-copaene content. Journal of Pest Science 89: 427–438. https://doi.org/10.1007/s10340-015-0708-5

Kirkendall LR. 2018. Invasive bark beetles (Coleoptera, Curculionidae, Scolytinae) in Chile and Argentina, including two species new for South America, and the correct identity of the Orthotomicus species in Chile and Argentina. Diversity 10: 40. https://doi:10.3390/d10020040

Kirkendall LR, Faccoli M. 2010. Bark beetles and pinhole borers (Curculionidae, Scolytinae, Platypodinae) alien to Europe. ZooKeys 56: 227–251. https://doi.org/10.3897/zookeys.56.529

Kirkendall LR, Ødegaard F. 2007. Ongoing invasions of old-growth tropical forests: establishment of three incestuous beetle species in Central America (Curculionidae, Scolytinae). Zootaxa 1588: 53–62. https://doi.org/10.11646/zootaxa.1588.1.3

Knížek M. 1988. Xyleborus alni Niijima, 1909. Acta Entomologica Bohemoslovaca 85: 396.

Knížek M. 2011. Scolytinae. In: Löbl I, Smetana A (Eds) Catalogue of Palaearctic Coleoptera Vol. 7, Curculionoidea I. Apollo Books, Stenstrup, 204–251.

Kolenati FA. 1846. Meletemata Entomologica. Fascicule III. Brachyelytra Caucasi cum distributione geographica adnexis Pselaphinis, Scydmaenis, Notoxibus, et Xylophagis. Petropoli: Typis Imperialis Academiae Scientarum [6] + 44 pp., pls 12–14.

Kôno M 1938. Neue und wenig bekannte Ipiden als Schädlinge an Sachalintannen und Ezofichten in Hokkaido. Insecta Matsumurana 12: 64–73.

Kurentzov AI. 1941. Koroedy Dalnego Vostoka SSSR [Bark-beetles of the Far East, USSR]. Moskva, Leningrad: Izdatelstvo Akademii Nauk SSSR, 234 pp.

Kurentzov AI. 1948. Novye dannye po faune koroedov (Coleoptera, Ipidae) Primorskogo Kraya [New data on the bark-beetle fauna of the Maritime Region (Soviet Far East). Entomologischeskoe Obozrenie 30: 50–52.


Landi L, Braccini CL, Knížek M, Pereyra VA, Marvaldi AE. 2019. A newly detected exotic ambrosia beetle in Argentina Euwallacea interjectus (Coleoptera: Curculionidae: Scolytinae). Florida Entomologist 102: 240–242. https://doi.org/10.1653/024.102.0141

Landi L, Gómez D, Braccini CL, Pereyra VA, Smith SM, Marvaldi AE. 2017. Morphological and molecular identification of the invasive Xylosandrus crassiusculus (Motschulsky) (Coleoptera: Curculionidae: Scolytinae) and its South American range extending into Argentina and Uruguay. Journal of Economic Entomology 110: 344–349. https://doi.org/10.1093/aesa/sax032

Le Pelley RH. 1968. Pests of coffee. London and Harlow, UK: Longmans, Green and Co. Ltd.

Lea AM. 1894. Description of new species of Bostrychidae. Proceedings of the Linnean Society of New South Wales 8: 317–323.

Lea AM. 1910. On Australian and Tasmanian Coleoptera, with descriptions of new species Part I. Proceedings of the Royal Society of Victoria, New Series 22: 113–152, pl. 30.

Li Y, Bateman CC, Skelton J, Jusino MA, Nolen ZJ, Simmons DR, Hulcr J. 2017. Wood decay fungus Flavodon ambrosius (Basidiomycota: Polyporales) is widely farmed by two genera of ambrosia beetles. Fungal Biology 121: 984–989. https://doi.org/10.1016/j.funbio.2017.08.004

Lin C-S, Smith SM. 2022 Four  new species and five new records of xyleborine ambrosia beetles (Coleoptera, Curculionidae, Scolytinae, Xyleborini) from Taiwan. Taiwania 67: 285–301. https://doi.org/10.6165/tai.2022.67.285

Lightle DM, Gandhi KJK, Cognato AI, Mosley BJ, Nielsen DG, Herms DA. 2007. New reports of exotic and native ambrosia and bark beetle species (Coleoptera: Curculionidae: Scolytinae) from Ohio. The Great Lakes Entomologist 40: 194–200.

Lin W, Li Y, Johnson AJ, Gao L. 2019. New area records and new hosts of Ambrosiodmus minor (Stebbing) (Coleoptera: Curculionidae: Scolytinae) in mainland China. The Coleopterists Bulletin 73: 684–686. https://doi.org/10.1649/0010-065X-73.3.684


Madoffe S, Bakke A. 1995. Seasonal fluctuations and diversity of bark and wood-boring beetles in lowland forest: implications for management practices. South African Forestry Journal 173: 9–15. https://doi.org/10.1080/00382167.1995.9629684

Maiti PK, Saha N. 1986. Contributions to the knowledge of the bark and timber beetles (Scolytidae: Coleoptera) of the Andaman and Nicobar Islands. Records of the Zoological Survey of India, Miscellaneous Publications, Occasional Papers 86: 1–182.

Maiti PK, Saha N. 2004. Fauna of India and the adjacent countries. Scolytidae: Coleoptera (bark and ambrosia beetles). Vol. 1. Part 1. Zoological Survey of India, Kolkata, 268 pp.

Mandelshtam MYu. 2001. New synonymy and new records in Palaearctic Scolytidae (Coleoptera). Zoosystematica Rossica 9: 203–204.

Mandelshtam MY. 2006. New synonymies and new combinations in Scolytidae from the Kuril Archipelago and continental territories of the Russian Far East (Coleoptera). Zoosystematica Rossica 15: 323–325.

Mandelshtam, M.Yu. and Petrov, A.V., 2009. Family Scolytidae—bark beetles, in Nasekomye Lazovskogo zapovednika (Insects of the Lazovsky Reserve), Vladivostok: Dal’nauka, pp. 206–209.

Mandelshtam MY, Nikitsky NB. 2010. Review of Scolytidae (Coleoptera) type specimens from V. Motschulsky collection preserved in the Zoological Museum of Moscow State University. Byulleten’ Moskovskogo Obshchestva Ispytatelei Prirody Otdel Biologicheskii 115(5): 13–21.

Mandelshtam M Yu, Petrov AV, Smith SM, Cognato AI. 2019. Resurrection of Heteroborips Reitter, 1913 (Coleoptera: Curculionidae: Scolytinae) from synonymy with Xyleborus Eichhoff, 1864. The Coleopterists Bulletin 73: 387−394. https://doi.org/10.1649/0010-065X-73.2.387

Mandelshtam M Yu, Yakushkin EA, Petrov AV. 2018. Oriental ambrosia beetles (Coleoptera: Curculionidae: Scolytinae): new inhabitants of Primorsky krai in Russia. Russian Journal of Biological Invasions 9(4): 355–365.  https://doi.org/10.1134/S2075111718040082

Maner ML, Hanula JL, Braman SK. 2013. Gallery productivity, emergence, and flight activity of the redbay ambrosia beetle (Coleoptera: Curculionidae: Scolytinae). Environmental Entomology 42: 642–647. https://doi.org/10.1603/EN13014

Marchioro M, Faccoli M, Dal Cortivo M, Branco M, Roques A, Garcia A, Ruzzier E. 2022. New species and new records of exotic Scolytinae (Coleoptera, Curculionidae) in Europe. Biodiversity Data Journal 10: e93995. https://doi.org/10.3897/BDJ.10.e93995

Markalas S, Kalapanida M. 1997. Flight pattern of some Scolytidae attracted to flight barrier traps baited with ethanol in oak forest in Greece. Anzeiger für Schädlingskunde, Pflanzenschutz, Umweltschutz 70: 55–67. https://doi.org/10.1007/BF01996922

Mayers CG, Harrington TC, Biedermann PHW. 2022. Mycangia define the diverse ambrosia beetlefungus symbioses (pp. 105-142). In:  Schultz TR, Gawne R, Peregrine PN, eds. The Convergent Evolution of Agriculture in Humans and Insects, The MIT Press, Cambridge, MA.

Mayfield III AE, MacKenzie M, Cannon P, Oak S, Horn S, Hwang J, Kendra PE. 2013. Suitability of California bay laurel and other species as hosts for the non-native redbay ambrosia beetle and granulate ambrosia beetle. Agricultural and Forest Entomology 15: 227–235. https://doi.org/10.1111/afe.12009

McPherson BA, Erbilgin N, Wood DL, Svihra P, Storer AJ, Standiford RB. 2008. Attraction of ambrosia and bark beetles to coast live oaks infected by Phytophthora ramorum. Agricultural and Forest Entomology 10: 315–321. https://doi.org/10.1111/j.1461-9563.2008.00386.x

Mendel Z, Protasov A, Maoz Y, Maymon M, Miller G, Elazar M, Freeman S. 2017. The role of Euwallacea nr. fornicatus (Coleoptera: Scolytinae) in the wilt syndrome of avocado trees in Israel. Phytoparasitica 45: 341–359. https://doi.org/10.1007/s12600-017-0598-6

Mendel Z, Protasov A, Sharon M, Zveibil A, Yehuda SB, O’Donnell K, Rabaglia R, Wysocki M, Freeman S. 2012. An Asian ambrosia beetle Euwallacea fornicatus and its novel symbiotic fungus Fusarium sp. pose a serious threat to Israeli avocado industry. Phytoparasitica 40: 235–238. https://doi.org/10.1007/s12600-012-0223-7

Mizuno T, Kajimura H. 2002. Reproduction of the ambrosia beetle, Xyleborus pfeili (Ratzeburg) (Col., Scolytidae), on semi-artificial diet. Journal of Applied Entomology 126: 455–462. https://doi.org/10.1046/j.1439-0418.2002.00691.x

Montrouzier X. 1861. Essai sur la faune entomologique de la Nouvelle Calédonie et des îles des Pins, Art, Lifu, etc. (suite). Annales de la Société Entomologique de France, série 4, 1: 265–306. 265–306.

Motschulsky V. 1863. Essai d'un catalogue des insectes de l'île Ceylan. (Suite). Bulletin de la Société Impériale des Naturalistes de Moscou 36: 421–532.

Motschulsky V. 1866. Essai d'un catalogue des insectes de l'île de Ceylan. (Supplément). Bulletin de la Société Impériale des Naturalistes de Moscou 39: 393–446.

Mulsant ME, Rey C. 1856. Description d'une nouvelle espèce de Coléoptère du genre Bostrichus. Opuscules Entomologiques 7(1856): 111–113. https://doi.org/10.5962/bhl.title.2682

Murayama J. 1930. Révisions des familes des Ipides et des Platypides de Corée. Journal of the Chosen Natural History Society 11: 1–34, 2 pls.

Murayama J. 1931. Révision des familles des Ipides et Platypides (Coléoptères) de l'ile de Quelpart. Annotationes Zoologicae Japonenses 13: 39–62, 2 pls.

Murayama J. 1934. Notes on the Ipidae (Coleoptera) from Kiushu. Annotationes Zoologicae Japonenses 14: 287–300.

Murayama J. 1936. Notes sur les Scolytides (Coléoptères) de Honshû et Kiushû, Japon. Tenthredo 1: 121–149.

Murayama J. 1943. Nouvelles espèces des Scolytides (Coléoptères) du Manchoukuo Annotationes Zoologicae Japonenses 22: 96–100.

Murayama J. 1950. A new genus and some new species of Scolytidae from Japan (Coleoptera). Transactions of the Shikoku Entomological Society 1: 49–53.

Murayama J. 1951. New genus and species of Scolytidae (Coleoptera) from Ohshima and Shionomisaki, Wakayama prefecture. Bulletin of the Faculty of Agriculture, Yamaguti University 2: 1–7.

Murayama J. 1952. Notes on the scolytid-beetles (Coleoptera) from southern and western parts of Izu Peninsula, Shizuoka Prefecture. Bulletin of the Faculty of Agriculture, Yamaguti University 3: 15–23.

Murayama J. 1953. The insect fauna of Mt Ishizuchi and Omogo valley, Iyo, Japan. The Scolytidae and Platypodidae (Coleoptera). Transactions of the Shikoku Entomological Society 3: 144–165.

Murayama J. 1954. Scolytid fauna on the northern half of Honshu with a distribution table of all the scolytid-species described from Japan. Bulletin of the Faculty of Agriculture, Yamaguti University 5: 149–212.

Murayama J. 1955. Supplementary notes on the scolytid-fauna of Japan. Bulletin of the Faculty of Agriculture, Yamaguti University 6: 81–106, pls. 3–4.

Murayama J. 1958. Studies in the scolytid-fauna of the northern half of the Far East, IV: new genera and new species. Bulletin of the Faculty of Agriculture, Yamaguti University 9: 927–936.

Murayama J. 1961. Scolytid-beetles from Niigata Prefecture, Japan. Akitu, Kyoto 10: 23–32.

Murayama J, Kalshoven LGE. 1962. Xyleborus morstatti Hag, a synonym of X. compactus Eichh. (Col, Scolytidae). Entomologische Berichten, Amsterdam 22: 247–250.

Murphy DH, Meepol W. 1990. Timber beetles of the Ranong mangrove forests. Mangrove Ecosystems Occasional Papers 7: 5–8.


Na F, Carrillo JD, Mayorquin JS, Ndinga-Muniania C, Stajich JE, Stouthamer R, Huang YT, Lin YT, Chen CY, Eskalen A. 2018. Two novel fungal symbionts Fusarium kuroshium sp. nov. and Graphium kuroshium sp. nov. of Kuroshio shot-hole borer (Euwallacea sp. nr. fornicatus) cause Fusarium dieback on woody host species in California. Plant Disease 102: 1154–1164. https://doi.org/10.1094/PDIS-07-17-1042-RE

Nakashima T, Otomo T, Owada Y, Iizuka T. 1992. SEM observations on growing conditions of the fungi in the galleries of several ambrosia beetles: (Coleoptera: Scolytidae and Platypodidae). Journal of the Faculty of Agriculture, Hokkaido University 65: 239–273.

Niisima Y. 1909. Die Scolytiden Hokkaidos unter Berücksichtigung ihrer Bedeutung für Forstschäden. Journal of the College of Agriculture, Tohoku Imperial University, Sapporo 3: 109–179.

Niisima Y. 1910. Die Borkenkaefer Nord- und Mittel-Japans. Transactions of the Sapporo Natural History Society 3: 1–18.

Nikulina T, Mandelshtam M, Petrov A, Nazarenko V, Yunakov N (2015) A survey of the weevils of Ukraine. Bark and ambrosia beetles (Coleoptera: Curculionidae: Platypodinae and Scolytinae) Zootaxa 3912: 1–61. http://dx.doi.org/10.11646/zootaxa.3912.1.1

Nobuchi A. 1979. Ambrosia beetles of mahogany in the Philippines (Coleoptera: Scolytidae and Platypodidae). Kontyû 47: 406–407.

Nobuchi A. 1966. Bark-beetles injurious to pine in Japan. Bulletin of the Government Forest Experiment Station,Tokyo 185: 1-49, pls. 1–6. (in Japanese; summary and key in English)

Nobuchi A. 1981a. Studies on Scolytidae (Coleoptera) XXII. Six new species and two new females of the genus Xyleborus from Japan. Kontyû 49: 143–154.

Nobuchi A. 1981b. Studies on Scolytidae XXIII. The ambrosia beetles of the genus Xylosandrus Reitter from Japan (Coleoptera). Bulletin of the Forestry and Forestry Products Research Institute 314: 27–37.

Nobuchi A. 1985. Family Scolytidae. Check-list of Coleoptera of Japan 30: 1–32.

Norris DM. 1976. Chemical interdependencies among Xyleborus spp. ambrosia beetles and their symbiotic microbes. Material und Organismen, Beiheft 3: 479–488.

Nunberg M. 1956. Zmiany nazw i synonimika niektórych korników (Coleoptera, Scolytidae) [Namensänderungen und Synonymie einiger Borkenkäfer]. Annales Zoologici 16: 207–214.

Nunberg M. 1959. Die Gattung Xyleborus Eichhoff (Coleoptera: Scolytidae) Ergänzungen, Berichtigungen und Erweiterung der Diagnosen. Beiträge zur Entomologie 9: 413–466.

Nunberg M. 1961. Zur Kenntnis der malayischen und aethiopischen Borken- und Kernkäferfauna (Col. Scolytidae und Platypodidae). The Annals and Magazine of Natural History, series 13, 3: 609–632. https://doi.org/10.1080/00222936008651066

Nunberg M. 1963. Die Gattung Xyleborus Eichhoff (Coleoptera, Scolytidae). Ergänzungen, Berichtigungen und Erweiterung der Diagnosen (II. Teil). Annales du Musée Royale du Congo Belge, Série in 8°, Sciences Zoologiques 115: 1–127.


O’Donnell K, Sink S, Libeskind-Hadas R, Hulcr J, Kasson MT, Ploetz RC, Konkol JL, Ploetz JN, Carillo D, Campbell A, Duncan RE, Liyanage PN, Eskalen A, Na F, Geiser DM, Bateman C, Freeman S, Mendel Z, Sharon M, Aoki T, Cossé AA, Rooney AP. 2015. Discordant phylogenies suggest repeated host shifts in the Fusarium-Euwallacea ambrosia beetle mutualism. Fungal Genetics and Biology 82: 277–290. https://doi.org/10.1016/j.fgb.2014.10.014

Ohno S, Yoshioka K, Yoneyama K, Nakazawa H. 1988. The Scolytidae and Platypodidae (Coleoptera) from Solomon Islands, found in logs at Nagoya port, I. Research Bulletin of the Plant Protection Service, Japan 24: 91–95.

Ohno S, Yoshioka K, Uchida N, Yoneyama K, Tsukamoto K. 1989. The Scolytidae and Platypodidae (Coleoptera) from Bismark Archipelago found in logs at Nagoya port. Research Bulletin of the Plant Protection Service, Japan 25: 59–69.

Ohno S. 1990. The Scolytidae and Platypodidae (Coleoptera) from Borneo found in logs at Nagoya port 1. Research Bulletin of the Plant Protection Service, Japan 26: 83–94.

Okins KE, Thomas MC. 2010. New North American record for Xyleborinus andrewesi (Coleoptera: Curculionidae: Scolytinae). Florida Entomologist 93: 133–134. https://doi.org/10.1653/024.093.0122

Olivier AG. 1800. Entomologie, ou historire naturelle des Insectes, avec leurs caractères génériques et spécifiques, leur description, leur synonymie, et leur figure enluminée. Coléoptères. Tome quatrième. Paris: de Lanneau, 519 pp., 72 pls. https://doi.org/10.5962/bhl.title.49479

Owens D, Cruz LF, Montgomery WS, Narvaez TI, Schnell EQ, Tabanca N, Duncan RE, Carrillo D, Kendra PE. 2018. Host range expansion and increasing damage potential of Euwallacea nr. fornicatus (Coleoptera: Curculionidae) in Florida. Florida Entomologist 101: 229–237. https://doi.org/10.1653/024.101.0212


Palm T. 1959. Die Holz- und Rinden-Käfer der süd- und mittelschwedischen Laubbäume. Opuscula Entomologica, Supplementum 16: 1–374.

Panzer GWF. 1793. Faunae insectorum Germanicae initia oder Deutschlands Insekten. Heft 8. Nürenberg: Felsecker, 24pp., 24 pls. https://doi.org/10.5962/bhl.title.15007

Papp T, de Beer ZW, Migliorini D, Nel WJ, Wingfield MJ. 2018. The polyphagous shot-hole borer (PSHB) and its fungal symbiont Fusarium euwallaceae: a new invasion in South Africa. Australasian Plant Pathology 47: 231–237.   https://doi.org/10.1007/s13313-018-0545-0

Park S, Smith SM, Cognato AI, Beaver RA. 2020. Catalogue of Korean Xyleborine ambrosia beetles (Coleoptera: Curculionidae) with seven new species. Journal of Asia-Pacific Biodiversity 13: 210–228.  https://doi.org/10.1016/j.japb.2020.01.002

Peck WD. 1817. On the insects which destroy the young branches of the pear-tree, and the leading shoot of the weymouth-pine. Massachusetts Agricultural Repository and Journal 4: 205–2011, pls 1–II.

Peer K, Taborsky M. 2004. Female ambrosia beetles adjust their offspring sex ratio according to outbreeding opportunities for their sons. Journal of Evolutionary Biology 17: 257–264. https://doi.org/10.1111/j.1420-9101.2003.00687.x

Peer K, Taborsky M. 2005. Outbreeding depression, but no inbreeding depression in haplodiploid ambrosia beetles with regular sibling mating. Evolution 59: 317–323. https://doi.org/10.1111/j.0014-3820.2005.tb00992.x

Peer K, Taborsky M. 2007. Delayed dispersal as a potential route to cooperative breeding in ambrosia beetles. Behavioral Ecology and Sociobiology 61: 729–739. https://doi.org/10.1007/s00265-006-0303-0

Peña JE, Weihman SW, McLean S, Cave RD, Carillo D, Duncan RE, Evans G, Krauth S, Thomas MC, Lu SS, Kendra PE, Roda AL. 2015. Predators and parasitoids associated with Scolytinae in Persea species (Laurales: Lauraceae) and other Lauraceae in Florida and Taiwan. Florida Entomologist 98: 903–910. https://doi.org/10.1653/024.098.0314

Pennacchio F, Roversi PF, Francardi V, Gatti E. 2003. Xylosandrus crassiusculus (Motschulsky) a bark beetle new to Europe (Coleoptera Scolytidae). Redia 86: 77-80.

Pfeffer A. 1944. Bemerkungen zur Arbeit von Hans Eggers: Zur Palearktischen Borkenkäferfauna. VIII. Borkenkäfer aus dem asiatischen Russland (Col: Ipidae). Arbeiten über Morphologische und Taxonomische Entomologie aus Berlin-Dahlem 11: 130–131.


Rabaglia RJ, Okins KE. 2011. Entomology section. Tri-ology 50 (3): 6–9.

Rabaglia RJ, Dole SA, Cognato AI. 2006. Review of America Xyleborina (Coleoptera: Curculionidae: Scolytinae) occurring north of Mexico, with an illustrated key. Annals of the Entomological Society of America 99: 1034–1056. https://doi.org/10.1603/0013- 8746(2006)99[1034:ROAXCC]2.0.CO;2

Rabaglia RJ, Vandenburg N, Acciavatti RE. 2009. First records of Anisandrus maiche Stark (Coleoptera: Curculionidae: Scolytinae) from North America. Zootaxa 2137: 23–28. https://doi.org/10.11646/zootaxa.2137.1.2

Rabaglia RJ, Knížek M, Johnson W. 2010. First records of Xyleborinus octiesdentatus (Murayama) (Coleoptera, Curculionidae, Scolytinae) from North America. ZooKeys 56: 219–226. https://doi.org/10.3897/zookeys.56.528

Rabaglia RJ, Smith SL, Rugman-Jones P, DiGirolomo M, Ewing C, Eskalen A. 2020. Establishment of a non-native xyleborine ambrosia beetle, Xyleborus monographus (Fabricius) (Coleoptera: Curculionidae: Scolytinae), new to North America in California. Zootaxa 4786: 269–276. https://doi.org/10.11646/zootaxa.4786.2.8

Rangel R, Pérez M, Sánchez S, Capello S. 2012. Population fluctuation of Xyleborus ferrugineus and X. affinis (Coleoptera: Curculionidae) in ecosystems of Tabasco, Mexico. Revista de Biología Tropical 60: 1577–1588. https://doi.org/10.15517/rbt.v60i4.2075

Ranger CM, Reding ME, Persad AB, Herms DA. 2010. Ability of stress-related volatiles to attract and induce attacks by Xylosandrus germanus and other ambrosia beetles (Coleoptera: Curculionidae, Scolytinae). Agricultural and Forest Entomology 12: 177–185. https://doi.org/10.1111/j.1461-9563.2009.00469.x

Ranger CM, Schultz PB, Frank SD, Chong JH, Reding ME. 2015. Non-native ambrosia beetles as opportunistic exploiters of living but weakened trees. PLoS ONE 10: e0131496. https://doi:10.1371/journal.pone.0131496

Ranger CM, Reding ME, Schultz PB, Oliver JB, Frank SD, Addesso KM, Chong JH, Sampson B, Werle C, Gill S, Krause C. 2016. Biology, ecology, and management of non-native ambrosia beetles (Coleoptera: Curculionidae: Scolytinae) in ornamental plant nurseries. Journal of Integrated Pest Management 7: 9. https://doi.org/10.1603/EN11299

Ratzeburg JTC. 1837. Die Forst-insekten: oder Abbildung und Beschreibung der in den Wäldern Preussens und der Nachbarstaaten als schädlich oder nützlich bekannt gewordenen Insekten. Erster Theil. Die Käfer. Nicolai, Berlin. X +4 + 202 pp., 21 pls.https://doi.org/10.5962/bhl.title.34392

Reding ME, Schultz PB, Ranger CM, Oliver JB. 2011. Optimizing ethanol-baited traps for monitoring damaging ambrosia beetles (Coleoptera: Curculionidae: Scolytinae) in ornamental nurseries. Journal of Economic Entomology 104: 2017–2024. https://doi.org/10.1603/EC11119

Reding ME, Ranger CM, Oliver JB, Schultz PB. 2013. Monitoring attack and flight activity of Xylosandrus spp. (Coleoptera: Curculionidae: Scolytinae): The influence of temperature on activity. Journal of Economic Entomology 106: 1780–1787. https://doi.org/10.1603/EC13134

Reitter E. 1913. Bestimmungs-Tabelle der Borkenkäfer (Scolytidae) aus Europa und den angrenzenden Ländern. Wiener Entomologische Zeitung, 32, Beiheft, 1–116. http://doi.org/10.3931/e-rara-68983

Rey C. 1883. [new taxa]. In: Eichhoff WJ. Les xylophages d’Europe. Avec des notes et additions concernant la faune gallo-rhénane. (Suite et fin). Revue d’Entomologie 2: 121–145.

Roberts H. 1977. Observations on the biology of some tropical rain forest Scolytidae (Coleoptera) from Fiji II. Subfamily Ipinae- tribe Xyleborini. Journal of Natural History 11: 251–272. https://doi.org/10.1080/00222937700770181


Saha N, Maiti PK. 1984. On a collection of scolytid beetles (Scolytidae: Coleoptera) from Sikkim, India. Records of the Zoological Survey of India 81(3–4): 1–8.

Saha N, Maiti PK. 1987. Description of hitherto unknown males of three species of scolytid beetles (Scolytidae: Coleoptera) from India. Bulletin of the Zoological Survey of India 8: 71–76.

Saha N, Maiti PK. 1996. Insecta: Coleoptera: Scolytidae. Fauna of West Bengal, 6B: 775–866.

Saha N, Maiti PK, Chakraborti S. 1992. On some species of Xylosandrus Reitter (Coleoptera: Scolytidae) from the sub-Himalayan West Bengal with description of a new species. Records of the Zoological Survey of India 91: 9–27.

Sahlberg CR 1836. Dissertatio entomologica insecta fennica enumerans, cujus particulam decimam partis secundae, cons. ampl. facult. philos. ad Imper. Univers. Alexandr. in Fennia, publico offert examini Carolus Reginaldus Sahlberg, respondente Alfredo Wacklin, Ostrobottniensi. In audit. philos. die 7 Maji 1836. Aboae: J. C. Frenckel, pp. 145–160.

Sampson FW. 1911. On two new wood-boring beetles (Ipidae). The Annals and Magazine of Natural History, series 8, 8: 381–384. https://doi.org/10.1080/00222931108693046

Sampson FW. 1912. Some new species of Ipidae and Platypodidae in the British Museum. The Annals and Magazine of Natural History, series 8, 10: 245–250. https://doi.org/10.1080/00222931208693228

Sampson FW. 1913. Some hitherto undescribed Ipidae and Platypodidae from India and Burma. The Annals and Magazine of Natural History, series 8, 12: 443–452. https://doi.org/10.1080/00222931308693422

Sampson FW. 1914. No. XVIII. — Coleoptera; Platypodidae and Ipidae from the Seychelles Islands. Transactions of the Linnean Society of London, Zoology, Second Series 16: 379–391. http://dx.doi.org/10.1111/j.1096-3642.1913.tb00155.x

Sampson FW. 1919. Notes on Platypodidae and Scolytidae collected by Mr. G. E. Bryant and others. The Annals and Magazine of Natural History, series 9, 4: 105–114. http://dx.doi.org/10.1080/00222932108632486

Sampson FW. 1921. Further notes on Platypodidae and Scolytidae collected by Mr. G. E. Bryant and others. The Annals and Magazine of Natural History, series 9, 7: 25–37. https://doi.org/10.1080/00222932108632486

Sampson FW. 1922. Previously undescribed Scolytidae and Platypodidae from the Indian area. The Annals and Magazine of Natural History, series 9, 10: 145–152. https://doi.org/10.1080/00222932208632753

Sampson FW. 1923. Previously undescribed Scolytidae and Platypodidae from the Indian area, Part II. The Annals and Magazine of Natural History, series 9, 11: 285–289. https://doi.org/10.1080/00222932308632858

Samuelson GA. 1981. A synopsis of Hawaiian Xyleborini (Coleoptera: Scolytidae). Pacific Insects 23: 50–92.

Saruhan H, Akyol I. 2012. Monitoring population density and fluctuations of Anisandrus dispar and Xyleborinus saxesenii (Coleoptera: Scolytinae, Curculionidae) in hazelnut orchards. African Journal of Biotechnology 11: 4202–4207. http://dx.doi.org/10.5897/AJB11.4185

Schaufuss CFC. 1897. Beitrag zur Käferfauna Madagascars III. Missions scientifiques de M Ch Alluaud aux îles Séchelles (1892) et à Diego-Suarez, Madagascar (1893) (Scolytidae et Platypodidae). Tijdschrift voor Entomologie 40: 209–225.

Schedl KE. 1931. Notes on the genus Xyleborus Eichh. The Annals and Magazine of Natural History, series 10, 8: 339–347. https://doi.org/10.1080/00222933108673402

Schedl KE. 1933. New Scolytidae from the Philippines. Philippine Journal of Science 51: 101–107.

Schedl KE. 1934a. Neue Borkenkäfer. Entomologische Blätter 30: 37–39.

Schedl KE. 1934b. Neue indomalayische Scolytidae. II. Beitrag zur Morphologie und Systematik der Scolytoidea. Entomologische Berichten, Amsterdam 9: 84–92.

Schedl KE. 1934c. Scolytidae and Platypodidae In: Winkler A (Ed) Catalogus Coleopterorum Regionis Palaearctica. (4 volumes). By author, Wien, 1632–1647.

Schedl KE. 1935a. Fauna Philippinensis (Platypodidae et Scolytidae) III. Philippine Journal of Science 56: 395–403.

Schedl KE. 1935b. New bark beetles and ambrosia-beetles (Col). Stylops 4: 270–276. https://doi.org/10.1111/j.1365-3113.1935.tb00659.x

Schedl KE. 1936a. Notes on Malayan Scolytidae and Platypodidae and descriptions of some new species. Journal of the Federated Malay States Museums 18: 1–18.

Schedl KE. 1936b. Scolytidae and Platypodidae. Contribution 35. The collection of the South Australian Museum. Records of the South Australian Museum 5: 513–535.

Schedl KE. 1936c. Scolytidae and Platypodidae: Fauna Philippinensis, IV. Philippine Journal of Science 60: 59–67.

Schedl KE. 1936d. Some new Scolytidae and Platypodidae from the Malay Peninsula. Journal of the Federated Malay States Museums 18: 19–35.

Schedl KE. 1937a. Scolytidae and Platypodidae. Fauna Borneensis. Part I. Sarawak Museum Journal 4: 543–552.

Schedl KE. 1937b. Scolytidae und Platypodidae. 45. Beitrag. Vereinschrift der Gesellschaft Luxemburger Naturfreunde 31: 15–17.

Schedl KE. 1937c. Scolytidae und Platypodidae-Zentral und südamerikanische Arten. Arquivos do Instituto de Biologia Vegetal, Rio de Janeiro 3: 155–170.

Schedl KE. 1938. Scolytidae und Platypodidae. 41. Beitrag zur Morphologie und Systematik der Scolytoidea. Mitteilungen aus dem Zoologischen Museum in Berlin 23: 459–464.

Schedl KE. 1939a. Malaysian Scolytidae and Platypodidae (IV). 57th contribution. Journal of the Federated Malay States Museums 18: 327–364.

Schedl KE. 1939b. Scolytidae and Platypodidae. 47. Beitrag zur Morphologie und Systematik der Scolytoidea. Tijdschrift voor Entomologie 82: 30–53.

Schedl KE. 1940. Scolytidae and Platypodidae. 61. Contribution to the morphology and taxonomy of the Scolytoidea. The Annals and Magazine of Natural History, series 11, 5: 433–442. https://doi.org/10.1080/00222934008527057

Schedl KE. 1941. 77th Contribution to the morphology and taxonomy of the Scolytoidea. Proceedings of the Hawaiian Entomological Society 11: 109–116.

Schedl KE. 1942a. Forschungberichte zur Scolytoiden-Fauna der Malayischen Halbinsel, V. 80. Beitrag zur Morphologie und Systematik der Scolytoidea. Kolonialforstliche Mitteilungen 5: 169–218.

Schedl KE. 1942b. Interessante und neue Scolytiden und Platypodiden aus der australischen Region. 79. Beitrag zur Morphologie und Systematik der Scolytoidea. Mitteilung der Münchner Entomologische Gesellschaft 32: 162–201.

Schedl KE. 1942c. Neue Scolytidae aus Java. 76. Beitrag zur Morphologie und Systematik der Scolytoidea. Tijdschrift voor Entomologie 85: 1–49.

Schedl KE. 1948. New species and records of Australian Scolytidae. Proceedings of the Royal Society of Queensland 60: 25–29.

Schedl KE. 1949. Neotropical Scolytoidea. I. 97th Contribution to the morphology and taxonomy of the Scolytoidea (Col). Revista Brasileira de Biologia 9: 261–284.

Schedl KE. 1950a. Fauna Fijiana (Scolytoidea). 94. Contribution to the morphology and taxonomy of the Scolytoidea. Occasional Papers of the Bernice P. Bishop Museum 20: 35–54.

Schedl KE. 1950b. Fauna Indo-Malayensis, II. 104. Contribution to the morphology and taxonomy of the Scolytoidea. The Annals and Magazine of Natural History, series 12, 3: 892–900. https://doi.org/10.1080/00222935008654721

Schedl KE. 1951a. Fauna Indomalayaensis, I. Tijdschrift voor Entomologie 93: 41–98. https://doi.org/10.1080/00222935008654721

Schedl KE. 1951b. Fauna Samoanus (Scolytoides), I. 109. Contribution. Bernice P. Bishop Museum Occasional Papers 20: 131–156.

Schedl KE. 1952a. Fauna Philippinensis, VIII. 123. Contribution to the morphology and taxonomy of the Scolytoidea. Philippine Journal of Science 80: 363–371.

Schedl KE. 1952b. Formosan Scolytoidea, I. lll. Contribution. Philippine Journal of Science 81: 61–65.

Schedl KE. 1952c. Scolytoides Congolais, IV. 132. Contribution to the morphology and taxonomy of the Scolytoidea. Bulletin - Institut Royal des Sciences Naturelles de Belgique 28: 1–12.

Schedl KE. 1952d. Zur synonymie der Borkenkäfer I. Entomologische Blätter 47/48: 158–164.

Schedl KE. 1953a. Bark and ambrosia-beetles from Indochina. 127. Contribution to the morphology and taxonomy of the Scolytoidea. Revue Francaise d'Entomologie 20: 123–130.

Schedl KE. 1953b. Fauna Indomalayensis.- III. Contribution 133 to the morphology and taxonomy of the Scolytoidea. The Annals and Magazine of Natural History, series 12, 6: 288–304. https://doi.org/10.1080/00222935308654424

Schedl KE. 1953c. Fauna Sinensis, I. Entomologische Blätter 49: 22–30.

Schedl KE. 1953d. New Scolytoidea. Queensland Museum Memoirs 13: 80–83.

Schedl KE. 1954a. Fauna Indomalayensis, IV: 141. Beitrag zur Morphologie und Systematik der Scolytoidea. The Philippine Journal of Science 83: 137–159.

Schedl KE. 1954b. Scolytoidea from the Gold Coast. I. (135. Contribution to the morphology and taxonomy of the Scolytoidea). Revue de Zoologie et de Botanique Africaines 50: 45‒88.

Schedl KE. 1955a. Borken- und Ambrosiakäfer aus den pazifischen Raum. 150. Beitrag zur Morphologie und Systematik der Scolytoidea. Entomologische Arbeiten aus dem Museum G. Frey 6: 277–310.

Schedl KE. 1955b. Fauna Sinensis, II. 152. Beitrag zur Morphologie und Systematik der Scolytoidea. Entomologische Blätter 51: 45–46.

Schedl KE. 1957. Scolytoidea nouveaux du Congo Belge. II. Mission R. Mayné – K. E. Schedl 1952. Annales du Musée Royale du Congo Belge, Série 8°, Sciences Zoologiques 56: 1–162.

Schedl KE. 1958a. A few new African Scolytidae in the British Museum. 168. Contribution to the morphology and taxonomy of the Scolytoidea. The Annals and Magazine of Natural History, series 13, 1: 557–560. https://doi.org/10.1080/00222935808650982

Schedl KE. 1958b. Bark and timber beetles from Malaya. Malayan Forester 21: 99–105.

Schedl KE. 1958c. Zur Synonymie der Borkenkäfer II. Tijdschrift voor Entomologie 101: 141–155.

Schedl KE. 1959. A checklist of the Scolytidae and Platypodidae (Coleoptera) of Ceylon with descriptions of new species and biological notes. Transactions of the Royal Entomological Society of London 111: 469–534. https://doi.org/10.1111/j.1365-2311.1959.tb02874.x

Schedl KE. 1960a. Synonymies of bark beetles (Scolytidae), IV. 174. Contribution to the morphology and taxonomy of the Scolytoidea. The Coleopterists Bulletin 14: 5–12.

Schedl KE. 1960b. Zur Synonymie der Borkenkäfer, V. 181. Beitrag zur Morphologie und Systematik der Scolytoidea. Entomologische Blätter 56: 103–112.

Schedl KE. 1962a. Zur Synonymie der Borkenkäfer VI. 203. Beitrag zur Morphologie und Systematik der Scolytoidea. Entomologische Blätter 58: 201–211.

Schedl KE. 1962b. Synonymies of bark beetles VII. 204. Contribution to the morphology and taxonomy of the Scolytoidea. Annals and Magazine of Natural History, series 13, 4: 697‒699. https://doi.org/10.1080/00222936108651195

Schedl KE. 1963a. Scolytidae und Platypodidae Afrikas. Band II. Familie Scolytidae (Fortsetzung), Unterfamilie Ipinae (Fortsetzung). Revista de Entomologia de Moçambique 5: 1–594.

Schedl KE. 1963b. Zur Synonymie der Borkenkäfer IX. 209. Beitrag zur Morphologie und Systematik der Scolytoidea. Entomologische Abhandlungen und Berichte aus dem Staatlichen Museum für Tierkunde in Dresden 28: 257–268.

Schedl KE. 1964a. Neue und interessante Scolytoidea von den Sunda-Inseln, Neu Guinea, und Australien. 202. Beitrag zur Morphologie und Systematic der Scolytoidea. Tijdschrift voor Entomologie 107: 297–306.

Schedl KE. 1964b. Scolytoidea from Borneo III. 185. Contribution to the morphology and taxonomy of the Scolytoidea. Reichenbachia 4: 241–254.

Schedl KE. 1964c. Zur Synonymie der Borkenkäfer XIV. 223. Beitrag zur Morphologie und Systematik der Scolytoidea. Reichenbachia 2: 303–317.

Schedl KE. 1964d. Zur Synonymie der Borkenkäfer XV. 228. Beitrag zur Morphologie und Systematik der Scolytoidea. Reichenbachia 3: 303–317.

Schedl KE. 1965. Scolytidae und Platypodidae aus dem Naturhistorika Riksmuseum in Stockholm. 235. Beitrag zur Morphologie und Systematik der Scolytoidea. Arkiv för Zoologi 18: 17–31.

Schedl KE. 1966a. Check list of the Scolytidae and Platypodidae from the Philippine Islands. 196. Contribution to the morphology and taxonomy of the Scolytoidea. Entomologische Abhandlungen 35: 1–122.

Schedl KE. 1966b. Pin-hole borers and bark-beetles (Scolytidae and Platypodidae) intercepted from imported logs in Japanese ports. 241. Contribution to the morphology and taxonomy of the Scolytoidea. Kontyû 34: 29–43.

Schedl KE. 1967. Zur Synonymie der Borkenkäfer, XVI. 242. Beitrag zur Morphologie und Systematik der Scolytoidea. Entomologisk Tidskrift 88: 146–163.

Schedl KE. 1969a. Bark-beetles and pin-hole borers (Scolytidae and Platypodidae) intercepted from imported logs in Japanese ports, III. 258. Contribution to the morphology and taxonomy of the Scolytoidea. Kontyû 37: 202–219.

Schedl KE. 1969b. Indian bark and timber beetles, V. 217. Contribution to the morphology and taxonomy of the Scolytoidea. Oriental Insects 3: 47–70. https://doi.org/10.1080/00305316.1969.10433895

Schedl KE. 1970a. Another collection of Scolytidae and Platypodidae of economic importance from the territory of Papua and New Guinea. 254. Contribution to the morphology and taxonomy of the Scolytoidea. Proceedings of the Linnean Society of New South Wales 94: 128–132.

Schedl KE. 1970b. Zur Synonymie der Borkenkäfer, XX. Annalen des Naturhistorisches Museum Wien 74: 221–231.

Schedl KE. 1971a. Coleoptera: Scolytidae and Platypodidae from Ceylon. Entomologica Scandinavica Supplementum 1: 274–285.

Schedl KE. 1971b. Indomalayan bark and timber beetles. 273. Contribution to the morphology and taxonomy of the Scolytoidea. Oriental Insects 5: 361–399. https://doi.org/10.1080/00305316.1971.10434023

Schedl KE. 1972a. New Scolytidae and Platypodidae from the Papuan subregion and Australia. 279. Contribution to the morphology and taxonomy of the Scolytoidea. Papua New Guinea Agricultural Journal 23: 61–72.

Schedl KE. 1972b. Zur Synonymie der Borkenkäfer XXI. 281. Beitrag zur Morphologie und Systematik der Scolytoidea. Entomologische Arbeiten aus dem Museum G. Frey 23: 150–161.

Schedl KE. 1973. New Scolytidae and Platypodidae from the Papuan subregion. 299. Contribution to the morphology and taxonomy of the Scolytoidea. Papua New Guinea Agricultural Journal 24: 87–97.

Schedl KE. 1974. Borken- und ambrosiakäfer aus Vietnam (IV). 298. Beitrag zur Morphologie und Systematik der Scolytoidea. Travaux du Museum d'Histoire Naturelle Gregoire Antipa, Bukarest 14: 261–266.

Schedl KE. 1975a. Bark and timber beetles of the Oriental region. 319. Contribution to the morphology and taxonomy of the Scolytoidea. Oriental Insects 9: 451–460. https://doi.org/10.1080/00305316.1975.10434514

Schedl KE. 1975b. Indian bark and timber beetles VI. 312. Contribution to the morphology and taxonomy of the Scolytoidea. Revue Suisse de Zoologie 82: 445–458. https://doi.org/10.5962/bhl.part.78268

Schedl KE. 1975c. New Scolytidae and Platypodidae from Papua and New Guinea, IV. 317. Contribution to the morphology and taxonomy of the Scolytoidea. Annalen des Naturhistorisches Museum, Wien 79: 337–399.

Schedl KE. 1975d. New Scolytidae and Platypodidae from Papua/New Guinea (Coleoptera). 315. Contribution to the morphology and taxonomy of the Scolytoidea. Reichenbachia 15: 215–232.

Schedl KE. 1975e. Zur Synonymie der Borkenkäfer, XXVI. 318. Beitrag zur Morphologie und Systematik der Scolytoidea. Zeitschrift der Arbeitsgemeinschaft Österreichischer Entomologen 27: 33–38.

Schedl KE. 1977. Some new bark beetles from the Indomalayan region. 332. Contribution to the morphology and taxonomy of the Scolytoidea. Oriental Insects 11: 490–504. https://doi.org/10.1080/00305316.1977.11090920

Schedl KE. 1979a. Bark and timber beetles from Australia. 326. Contribution. Entomologischen Arbeiten aus dem Museum G. Frey 28: 157–164.

Schedl KE. 1979b. New Scolytidae and Platypodidae from Papua New Guinea (V). 311th Contribution to the morphology and taxonomy of the Scolytoidea. Faunistische Abhandlungen, Staatlichen Museum für Tierkunde in Dresden 7: 95–120.

Schedl KE. 1980. Zur synonymie der Borkenkäfer XXVIII. 339. Beitrag zur Morphologie und Systematik der Scolytoidea. Zeitschrift der Arbeitsgemeinschaft Österreichischer Entomologen 31: 117–124.

Schiefer TL, Bright DE. 2004. Xylosandrus mutilatus (Blandford), an exotic ambrosia beetle (Coleoptera: Curculionidae: Scolytinae: Xyleborini) new to North America. The Coleopterists Bulletin 58: 431–438. https://doi.org/10.1649/760

Schiefer TL. 2018. First record of the introduced ambrosia beetle Ambrosiophilus nodulosus (Eggers) in Mississippi, with notes on the distribution of Ambrosiodmus minor (Stebbing) (Coleoptera: Curculionidae: Scolytinae). The Coleopterists Bulletin 72: 384–385. https://doi.org/10.1649/0010-065X-72.2.384

Schneider I. 1987. Verbreitung, Pilzübertragung, und Brutsystem des Ambrosiakäfers Xyleborus affinis im vergleich mit X. mascarensis (Coleoptera: Scolytidae) Entomologia Generalis 12: 267–275. https://doi.org/10.1127/entom.gen/12/1987/267

Sharp D. 1885. In: Blackburn T & Sharp D: Memoirs on the Coleoptera of the Hawaiian Islands. The Scientific Transactions of the Royal Dublin Society 3: 119–300, pls. IV–V.

Sittichaya W. 2012. Bark and ambrosia beetles (Coleoptera: Curculionidae: Scolytinae and Platypodinae) infesting mango trees (Mangifera indica L.) in Southern Thailand, with new species recorded for Thailand. Songklanakarin Journal of Science and Technology 34: 153–155.

Sittichaya W, Beaver RA. 2018. Cnestus quadrispinosus, a new species of xyleborine ambrosia beetle from Thailand and Borneo (Coleoptera, Curculionidae, Scolytinae, Xyleborini). ZooKeys 795: 31–37. https://doi.org/10.3897/zookeys.795.28384

Sittichaya W, Smith SM, Beaver RA. 2019. Ten newly recorded species of xyleborine ambrosia beetles (Coleoptera, Curculionidae, Scolytinae, Xyleborini) from Thailand. ZooKeys 862: 109–127. https://doi.org/10.3897/ zookeys.862.34766

Sittichaya W, Smith SM. 2020. Eggersanthus Sittichaya and Smith gen. nov., a new Oriental ambrosia beetle genus (Coleoptera: Curculionidae: Scolytinae: Xyleborini). Zootaxa 4779: 282–288. http://dx.doi.org/10.11646/zootaxa.4779.2.9

Sittichaya W, Smith SM (2022) New species and new records of Amasa Lea, 1894 ambrosia beetles from Thailand (Coleoptera: Curculionidae: Scolytinae: Xyleborini). Zootaxa. In press.

Sittichaya W, Smith SM, Beaver RA, Thaochan N (2021) Revision of the xyleborine ambrosia beetle genus Microperus Wood, 1980 (Curculionidae: Scolytinae, Xyleborini) of Thailand; with four new species and four newly recorded species. ZooKeys 1074: 191–214. https://doi.org/10.3897/zookeys.1074.76235

Sivapalan P. 1975. The dispersion of brood galleries of Xyleborus fornicatus Eichh (Coleoptera, Scolytidae) in tea plants. Bulletin of Entomological Research 65: 501–506. https://doi.org/10.1017/S0007485300006167

Sivapalan P. 1977. Population dynamics of Xyleborus fornicatus Eichhoff (Coleoptera: Scolytidae) in relation to yield trends in tea. Bulletin of Entomological Research 67: 329–335. https://doi.org/10.1017/S0007485300011159

Six DL, Stone WD, de Beer ZW, Woolfolk SW. 2009. Ambrosiella beaveri sp. nov., associated with an exotic ambrosia beetle, Xylosandrus mutilatus (Coleoptera: Curculionidae, Scolytinae) in Mississippi, USA. Antonie van Leeuwenhoek 96: 17–29. https://doi.org/10.1007/s10482-009-9331-x

Smith SM, Cognato AI. 2015. Ambrosiophilus peregrinus n. sp., an exotic ambrosia beetle discovered in Georgia, U.S.A. (Coleoptera: Curculionidae: Scolytinae). The Coleopterists Bulletin 69: 213–220. https://doi.org/10.1649/0010-065X-69.2.213

Smith SM, Cognato AI. 2022. New exotic beetles found in the United States among pseudocryptic Cyclorhipidion species (Coleoptera: Curculionidae: Scolytinae: Xyleborini) revealed via a multigene phylogeny. Insect Systematics and Diversity Insect Systematics and Diversity 6(4), 2. https://doi.org/10.1093/isd/ixac014

Smith SM, Hulcr J. 2015. Scolytus and other economically important bark and ambrosia beetles. In: Vega FE, Hofstetter RW (Eds) Bark Beetles: Biology and ecology of native and invasive species. Elsevier, San Diego, CA, 495−531.

Smith SM, Beaver RA, Cognato AI. 2017. The ambrosia beetle Ambrosiophilus peregrinus, introduced to the United States, is Ambrosiophilus nodulosus (n. comb.) (Curculionidae: Scolytinae). The Coleopterists Bulletin 71: 552–553. https://doi.org/10.1649/0010-065X-71.3.552

Smith SM, Beaver RA, Cognato AI. 2018b. New synonymy, new combinations, and taxonomic changes in Japanese xyleborine ambrosia beetles (Coleoptera: Curculionidae: Scolytinae). Zootaxa 4521: 391–403. https://dx.doi.org/10.11646/zootaxa.4521.3.5

Smith SM, Gomez DF, Beaver RA, Hulcr J, Cognato, AI. 2019. Reassessment of the species in the Euwallacea fornicatus (Coleoptera: Curculionidae: Scolytinae) complex after the rediscovery of the “lost” type specimen. Insects 10: 261. https://doi:10.3390/insects10090261

Smith SM, Rabaglia RJ, Beaver RA, Thu PQ, Cognato AI. 2018a. Attraction of ambrosia beetles (Curculionidae: Scolytinae) to semiochemicals in Vietnam with new records and a new species. The Coleopterists Bulletin 72: 838–844. https://doi.org/10.1649/0010-065X-72.4.838

Smith SM, Beaver RA, Singh S, and Cognato AI. 2018c. Taxonomic clarification and neotype designation for three Indian xyleborine species (Coleoptera: Curculionidae: Scolytinae). Zootaxa 4394: 138–140. https://doi.org/10.11646/zootaxa.4394.1.9

Smith SM, Beaver RA, and Cognato AI. 2020a. Taxonomic changes for Indo-Malayan ambrosia beetles (Coleoptera: Curculionidae: Scolytinae: Xyleborini). The Coleopterists Bulletin 74: 37–40. https://doi.org/10.1649/0010-065X-74.1.37

Smith SM, Beaver RA, and Cognato AI. 2020b. A monograph of the Xyleborini (Coleoptera, Curculionidae, Scolytinae) of the Indochinese Peninsula (except Malaysia) and China. ZooKeys 983: 1-442. https://doi.org/10.3897/zookeys.983.52630

Smith SM, Urvois T, Roques A, Cognato AI. 2022a. Recognition of the pseudocryptic species Xylosandrus declivigranulatus (Schedl) as distinct from X. crassiusculus (Motschulusky) (Curculionidae: Scolytinae: Xyleborini). The Coleopterists Bulletin 76: 367–374. https://doi.org/10.1649/0010-065X-76.3.367

Smith SM, Beaver RA, Sittichaya W, Cognato AI. 2022b. One hundred eighteen taxonomic changes among Xyleborine ambrosia beetles (Coleoptera: Curculionidae: Scolytinae). Zootaxa Zootaxa 5194: 151–175. ttps://doi.org/10.11646/zootaxa.5194.2.1

Smith, SM, Beaver RA, Pham HT, Cognato AI. 2022c. New species and new records of Xyleborini from the Oriental region, Japan and Papua New Guinea (Coleoptera: Curculionidae: Scolytinae). Zootaxa. In press.

Speranza S, Bucini D, Paparatti B. 2009. New observation on biology of European shot-hole borer [Xyleborus dispar (F.)] on hazel in northern Latium (Central Italy). Acta Horticulturae 845: 539–542. https://doi.org/10.17660/ActaHortic.2009.845.84

Sreedharan K, Balakrishnan M, Sanuel S, Bhat P. 1991. A note on the association of wood-boring beetles and a fungus with the death of silver oak trees on coffee plantations. Journal of Coffee Research 21: 145–148.

Stebbing EP. 1907. On some Assam sal (Shorea robusta) insect pests, with notes upon some insects predaceous and parasitic upon them. Indian Forest Bulletin 11: 1–66.

Stebbing EP. 1908. On some undescribed Scolytidae of economic importance from the Indian Region I. Indian Forest Memoirs 1: 1–12.

Stebbing EP. 1909. On some undescribed Scolytidae of economic importance from the Indian Region, II. Indian Forest Memoirs 1: 13–32.

Stebbing EP. 1914. Indian forest insects of economic importance. Eyre & Spottiswoode, London, 648 pp. http://dx.doi.org/10.5962/bhl.title.9203

Stone WD, Nebeker TE. 2007. Distribution and seasonal abundance of Xylosandrus mutilatus (Coleoptera: Curculionidae). Journal of Entomological Science 42: 409–412. https://doi.org/10.18474/0749-8004-42.3.409

Stone WD, Nebeker TE, Gerard PD. 2007. Host plants of Xylosandrus mutilatus in Mississippi. Florida Entomologist 90: 191–195. https://doi.org/10.1653/0015-4040(2007)90[191:HPOXMI]2.0.CO;2

Storer CG, Breinholt JW, Hulcr J. 2015. Wallacellus is Euwallacea: molecular phylogenetics settles generic relationships (Coleoptera: Curculionidae: Scolytinae: Xyleborini). Zootaxa 3974: 391–400. https://doi.org/10.11646/zootaxa.3974.3.6

Stouthamer R, Rugman-Jones P, Thu PQ, Eskalen A, Thibault T, Hulcr J, Wang L-J, Jordal BH, Chen C-Y, Cooperband M, Lin C-S, Kamata N, Lu S-S, Masuya H, Mendel Z, Rabaglia R, Sanguansub S, Shih H-H, Sittichaya W, Zong S. 2017. Tracing the origin of a cryptic invader: phylogeography of the Euwallacea fornicatus (Coleoptera: Curculionidae: Scolytinae) species complex. Agricultural and Forest Entomology 19: 366–375. https://doi.org/10.1111/ afe.12215

Strohmeyer H. 1910. Ueber Kaffeeschädlinge auf der Insel Java. Entomologische Blätter 6: 186–187.

Strohmeyer H. 1912. H. Sauter's Formosa-Ausbeute. Ipidae und Platypodidae. Entomologische Mitteilungen 1: 38–42. https://doi.org/10.5962/bhl.part.25902

Swaine JM. 1918. Canadian bark beetles. Part II. A preliminary classification, with an account of the habits and means of control. Dominion of Canada Department of Agriculture, Entomological Branch, Technical Bulletin 14: 143 pp. https://doi.org/10.5962/bhl.title.27867

Swaine JM. 1934. Three new species of Scolytidae (Coleoptera). The Canadian Entomologist 66: 204–206. https://doi.org/10.4039/Ent66204-9


Tang W-Q. 2000. Biological characteristics of Xyleborus mutilatus and its control. Journal of Zhejiang Forestry College 17: 417–420. (Chinese with English abstract)

Terekhova VV, Skrylnik YuYe. 2012. Biological peculiarities of the alien for Europe Anisandrus maiche Stark (Coleoptera: Curculionidae: Scolytinae) bark beetle in Ukraine. Russian Journal of Biological Invasions 3: 139–144. https://doi.org/10.1134/S2075111712020105

Thurston GS, Slater A, Nei I, Roberts J, McLachlan Hamilton K, Sweeney JD, Kimoto T. 2022. New Canadian and provincial records of Coleoptera resulting from annual Canadian Food Inspection Agency surveillance for detection of non-native, potentially invasive forest insects. Insects 13: 708. https://doi.org/10.3390/insects13080708


Vandenberg NJ, Rabaglia RJ, Bright DE. 2000. New records of two Xyleborus (Coleoptera: Scolytidae) in North America. Proceedings of the Entomological Society of Washington 102: 62–68.

Veen H. 1897. Description of a new species of the genus Tomicus. Notes from the Leyden Museum 19: 135–136.


Walgama RS. 2012. Ecology and integrated pest management of Xyleborus fornicatus (Coleoptera: Scolytidae) in Sri Lanka. Journal of Integrated Pest Management 3: A1–A8. https://doi.org/10.1603/IPM11031

Walker F. 1859. Characters of some apparently undescribed Ceylon insects. The Annals and Magazine of Natural History, series 3, 3: 258–265.

Wang J, Smith SM, and Cognato AI. 2020. Immanus virago, a new species from Borneo (Coleoptera, Curculionidae: Scolytinae: Xyleborini). The Coleopterists Bulletin 74: 438–440. https://doi.org/10.1649/0010-065X-74.2.438

Weber BC, McPherson JE. 1983a. Life history of the ambrosia beetle Xylosandrus germanus (Coleoptera: Scolytidae). Annals of the Entomological Society of America 76: 455–462. https://doi.org/10.1093/aesa/76.3.455

Weber BC, McPherson JE. 1983b. World list of host plants of Xylosandrus germanus (Coleoptera: Scolytidae). The Coleopterists Bulletin 37: 114–134.

Wichmann HE. 1914. XXX Coleoptera, VII: Ipiden und Platypodiden In: Zoological results of the Abor Expedition, 1911-1912. Records of the Indian Museum 8: 411–414. https://doi.org/10.5962/bhl.part.1194

Wollaston TV. 1854. Insecta Maderensia; being an account of the insects of the Islands of the Madeiran group. John van Voorst, London, xliii + 634 pp., 13 pls. https://doi.org/10.5962/bhl.title.140700

Wollaston TV. 1857. Catalogue of the Coleopterous insects of Madeira in the collection of the British Museum, London. 234 p., 1 pl. http://dx.doi.org/10.5962/bhl.title.9900

Wollaston TV. 1867. Coleoptera Hesperidum; being an enumeration of the Coleopterous insects of the Cape Verde archipelago. J. van Voorst, London xxxix + 285 pp., 1 map. https://doi.org/10.5962/bhl.title.48651

Wood SL. 1957. Distributional notes on and synonymies of some North American Scolytidae (Coleoptera). The Canadian Entomologist 89: 396–403. https://doi.org/10.4039/Ent89396-9

Wood SL. 1960. Coleoptera: Platypodidae and Scolytidae. Insects of Micronesia 18: 1–73.

Wood SL. 1962. Miscellaneous taxonomic notes on Scolytidae (Coleoptera). The Great Basin Naturalist 21: 87–107. https://doi.org/10.5962/bhl.part.11210

Wood SL. 1969. New synonymy and records of Platypodidae and Scolytidae (Coleoptera). The Great Basin Naturalist 29: 113–128. https://doi.org/10.5962/bhl.part.17054

Wood SL. 1972. New synonymy in American bark beetles (Scolytidae: Coleoptera), Part II. Great Basin Naturalist 32: 190–201. https://doi.org/10.5962/bhl.part.25774

Wood SL. 1974. New synonymy and records of American bark beetles (Coleoptera: Scolytidae). The Great Basin Naturalist 34: 277–290. https://doi.org/10.5962/bhl.part.15523

Wood SL. 1975a. New synonymy and new species of American bark beetles (Coleoptera: Scolytidae). The Great Basin Naturalist 35: 21–32.

Wood SL. 1975b. New synonymy and new species of American bark beetles (Coleoptera: Scolytidae) Part II. The Great Basin Naturalist 35: 391–401.

Wood SL. 1977. Introduced and exported American Scolytidae (Coleoptera). The Great Basin Naturalist 37: 67–74.    

Wood SL. 1980. New genera and new generic synonymy in Scolytidae (Coleoptera). The Great Basin Naturalist 40: 89– 97. https://doi.org/10.5962/bhl.part.20006

Wood SL. 1982. The bark and ambrosia beetles of North and Central America (Coleoptera: Scolytidae), a taxonomic monograph. Great Basin Naturalist Memoirs 8: 1–1359.

Wood SL. 1983. New synonymy and new species of American bark beetles (Coleoptera: Scolytidae), Part IX. The Great Basin Naturalist 43: 647–659. https://doi.org/10.5962/bhl.part.4489

Wood SL. 1984. New generic synonymy and new genera of Scolytidae (Coleoptera). The Great Basin Naturalist 44: 223–230.

Wood SL. 1986. A reclassification of the genera of Scolytidae (Coleoptera). Great Basin Naturalist Memoirs 10: 1–126.

Wood SL. 1989. Nomenclatural changes and new species of Scolytidae (Coleoptera), part IV. The Great Basin Naturalist 49: 167–185. https://doi.org/10.5962/bhl.part.22642

Wood SL. 1992. Nomenclatural changes and new species of Platypodidae and Scolytidae (Coleoptera), part II. The Great Basin Naturalist 52: 78–88.

Wood SL. 2007. Bark and ambrosia beetles of South America (Coleoptera: Scolytidae). Brigham Young University, M.L. Bean Life Science Museum; Provo, 900 p.

Wood SL, Bright DE. 1992. A catalog of Scolytidae and Platypodidae (Coleoptera), Part 2: Taxonomic index. The Great Basin Naturalist Memoirs 13: 1–1533.

Wurth T. 1908. De boeboek (Xyleborus coffeae n. sp.) op Coffea robusta. Mededeelingen van het Algemeen-Proefstation op Java te Salatiga (2) 3: 1–20, pls. i–iii.


Yin H-F, Huang F-S, Li Z-L. 1984. Economic insect fauna of China. Fasc. 29. Coleoptera: Scolytidae. Science Press, Beijing, x + 205 pp., XVII Tab. (Chinese with English abstract)


Zheng S, Li Y, Wang S. 2017. Identification and distribution of Xyleborinus artestriatus Eichhoff 1878 (Coleoptera: Curculionidae: Scolytinae) in China. Journal of Environmental Entomology 40: 1445–1450. (Chinese with English abstract).

Zimmermann C. 1868. Synopsis of the Scolytidae of America north of Mexico. Transactions of the American Entomological Society 2: 141–149. https://doi.org/10.2307/25076203