Lepidoptera is one of the largest and most economically important orders of plant eating insects. More than 6,000 species are
considered agricultural pests, with the largest number of pests in the superfamilies Noctuoidea, Pyraloidea, and Tortricoidea. Thousands of lepidopterans
are intercepted at U.S. ports of entry each year, and all life stages are encountered; however, the overwhelming majority of interceptions are larvae.
Identification of these specimens is essential to prevent the introduction of damaging agricultural pests into the U.S.
Although a large number of Lepidoptera are intercepted at U.S. ports of entry, relatively few species make up the majority of
named records. Thirty-five taxa listed in the USDA's PestID database (see list below) comprise nearly 80% of the total number of larval interceptions
in the past ten years; e.g., positive identification of only 35 taxa results in an 80% success rate for Lepidoptera larvae that
are currently being identified.
LepIntercept is a comprehensive identification resource for intercepted Lepidoptera larvae. It is specifically designed to be used
by identifiers working at U.S. ports of entry, although the information contained in the fact sheets and keys will be valuable to any
lepidopterist working with larvae and/or exotic pests. LepIntercept includes detailed fact sheets
treating the most frequently intercepted Lepidoptera larvae, an interactive identification key for these taxa, a comprehensive dichotomous key
treating all intercepted Lepidoptera, smaller dichotomous keys for various genera, and more than 300 photographs of diagnostic larval characters.
Each fact sheet contains information on taxonomy, host/origin, recorded distribution, and a highly detailed larval diagnosis and
identification authority discussion. In addition, detailed setal map illustrations are provided for all taxa; these are the first
complete setal maps published for many of these species.
The following species are treated in LepIntercept (ordered by number of interception records). *Non-Rep* denotes that the species is
non-reportable according to USDA-APHIS guidelines and is permissible when found entering the country.
* = Not ranked in the top 35 but included with Spodoptera litura
How to Use LepIntercept
LepIntercept includes two types of larval keys: traditional dichotomous keys and an interactive Lucid key. The keys are designed to complement the information on the fact
sheets in providing identification of Lepidoptera larvae frequently intercepted at U.S. ports of entry. Specific use of the keys is detailed below.
The LepIntercept Dichotomous Keys are traditional dichotomous keys, written with contrasting couplets, that are for advanced users.
A PDF version of each key is provided for downloading and/or printing.
Each key incorporates detailed morphological characters along with origin and host data (in most instances) to provide an identification. The level of identification may be to
a particular species, species group, genus, or higher taxonomic category. Not all taxa included in the dichotomous keys are specifically treated in LepIntercept, although
most are discussed in relation to other taxa that are treated. Unless specifically stated otherwise, all taxa are assumed to be late instar larvae.
The identifier is successful when he or she has navigated through a key to an ending couplet.
Users of the dichotomous keys should consult the detailed information provided in the LepIntercept fact
sheets (for those species treated) to verify identifications. In some instances, information in the keys is not sufficient to separate closely related species
(such as H. armigera from H. zea) and DNA data or other resources may be necessary to provide or confirm identifications for many intercepted larvae.
Lucid Interactive Identification Key
The LepIntercept Interactive Identification Key is written in Lucid 3.5 and must be run from the LepIntercept website. This key is for
more novice or casual users, and provides illustrations of morphological character states along with origin (and limited host) information to produce an identification. Only taxa treated in
LepIntercept are included in the key, and, in most cases, the Lucid key is not designed to separate closely related taxa, but rather to point the user to the appropriate
family, subfamily, or genus. In all cases, the user should consult the appropriate fact sheets for taxa remaining in the key. The key starts by forcing the user to choose the
origin of the larva (or unknown if the user does not know or is uncertain - do not guess origin!). The key then prompts for the user to select the state of the
prespiracular group on T1 (macro- vs. microlepidoptera), followed by other specific characters within these two general categories. Superfamily and family-specific
characters are displayed once the user has navigated the preliminary categories. Taxa are scored assuming a late instar larva in all instances. Host choices under each family or superfamily
represent the most common host-species combinations for a particular group - although useful, this information should be used with caution because many of species treated in
LepIntercept are polyphagous and cannot be diagnosed using only host. Consult the fact sheets for a discussion of potential hosts and origins in all cases. The complete character matrix for
the interactive key is available below as a PDF.
The identifier is successful when he or she has scored as many characters states as possible and one or more entities remain. In most instances, information in the key is not
sufficient to separate closely related species (such as H. armigera from H. zea). The identifier should consult the fact sheets for any remaining entities
(next to the species name) to determine if the description of the remaining taxa are consistent with the specimen being examined. The dichotomous keys can be used to obtain a more
definitive identification if the user is comfortable with larval morphology and navigating complicated couplets. DNA data may be necessary to provide or confirm identifications
for many intercepted larvae.
Detailed fact sheets for each taxon are provided under the Fact Sheets menu item. Fact sheets are
arranged alphabetically by family and subfamily. Fact sheet information is organized under tabs that contain the following sections: Overview, Detailed Information,
Interception Records, and Setal Map(s). Details on the information found in each of these sections is presented below. Because different web
browsers result in printing inconsistencies, a printable/downloadable PDF is available at the bottom of the first page (Overview) of each fact sheet.
The Overview tab contains a quick overview of all of the information for a particular taxon. This page contains the following sections:
- Taxonomy: contains information on the most current taxonomy for each taxon. The first line lists the superfamily, family, subfamily, tribe (for some taxa), genus,
species, and author (for species-level taxa). English common names and synonyms are listed for many taxa. Other notes provide information about the taxonomy of a particular
- Larval diagnosis (Summary): a summary of characters (usually morphological) that is used to diagnose each taxon is provided here. Nearly all morphological characters are
illustrated with one or more figures. A detailed discussion of these characters is provided in the Detailed Information section.
- Host/origin information: a summary of the host and origin information for each taxon from the PestID database is presented here. In many instances this information
is useful in diagnosing larval specimens, especially when a particular taxon is consistently encountered on the same host/origin combination.
- Recorded distribution: the recorded distribution for a particular taxon is provided here.
- Identification authority (Summary): a summary of the level of identification (family, genus, species) that should be attempted for interceptions of particular taxa from a
particular zoogeographic region at U.S. ports of entry. We did not compare exotic pests to possible non-targets from North America.
- Pest characterization: this is the ranking of a particular taxon using the criteria outlined by Cavey (2001). Information on the taxonomy, distribution, and
potential impact of each taxon is used to determine if it is quarantine significant for the U.S.
The Detailed Information tab contains a detailed discussion of the larval diagnosis, identification authority, and taxonomy (for some taxa). This page contains the
- Larval diagnosis (Detailed): a detailed discussion of the diagnosis for a particular taxon. Includes a review of literature descriptions and illustrations,
a list of taxonomically important characters provided by different authors, and discussion of these characters in relation to the identification of larvae intercepted
at U.S. ports of entry.
- Identification authority (Detailed): provides detailed information on the level of identification (family, genus, species) that should be attempted for
interceptions of particular taxa from a particular zoogeographic region at U.S. ports of entry. Information includes discussion of origin, host, morphological
characters, and/or related species. Anything could theoretically come from anywhere; we did not assume trans-shipping for any of the identification authorities.
Nor did we account for changes in foreign ports. Authorities may change and need modification as new trade patterns develop.
- Taxonomy (Detailed): (for some taxa), a detailed discussion of recent taxonomic changes. Included when recent taxonomic changes might confuse users who are used to referring
to a pest species in another genus.
A list of the interception records for a particular taxon from the PestID database are listed here:
- Origin records: a list of origins that are recorded for a particular taxon.
- Host records: a list of hosts for a particular taxon. Plant names are listed as exported from PestID and may be taxonomically incorrect in some instances.
A complete setal map is provided for each taxon (with the exception of Strymon because of the absence of primary setae). Setal map illustrations
were produced using Adobe Illustrator and slides of larval skins and/or detailed photographs of larval characters. A link to download a printable PDF of the setal
map is provided under each illustration.
Figures in the fact sheets were photographed by the authors of LepIntercept unless otherwise stated. All figures are available in two sizes.
Clicking on the small images on the right side of the page will display a much larger image in the center of the page. Large images range in size from 2400px wide to
1000px wide but are designed to scale to the size of the web browser window. Large images have a set of controls at the bottom of each image;
these have the functions outlined here:
Move - move the image in the browser window
Enlarge (if possible) - enlarge the image to full size
Close - close the image (can also simply click on the image again to close)
Advance (back) - move to the previous image
Play (slideshow) - display the images on the page in order
Advance (forward) - move to the next image
In LepIntercept you will find an interactive key designed to aid in the identification of
Lepidoptera intercepted at U.S. ports of entry. The interactive key was developed using Lucid technology.
Lucid software for creating and using interactive identification
keys was developed by the Centre
for Biological Information Technology at the University of Queensland
in Australia. Visit the Lucidcentral
website for more information on Lucid.
The interactive Lucid key runs as a Lucid Key Server application. Use of the key (and the rest of the
Internet Explorer, Firefox, Chrome, or Safari. LepIntercept has been successfully tested on the following
- Internet Explorer 9.X, 10.X, 11.X
- Firefox 23.X, 26.X
- Chrome 31.X
- Opera 15.X
The main LepIntercept window is 960px wide and is optimized to display best on screens
1024px or wider. This resolution covers all modern computer monitors as well as many portable devices.
Most tablet computers and smart phones are able to view the web pages and keys in "normal" browser mode.
NOTE: Some web pages, such as fact sheets
attached to items in Lucid interactive keys, may be considered pop-ups
by certain web browsers. If your browser or an add-on application is set to block
pop-ups, you may not be able to view all of the content on these pages. Additionally,
some web browsers or add-on applications may block "active content" on web
pages or interactive keys. To take full advantage of the information presented in this
Lucid tool, please consult your web browser or add-on application's help file for
assistance in disabling pop-up blockers and allowing active content.
LepIntercept was developed under a cooperative agreement between the authors, Colorado State University (CSU),
and the USDA-APHIS-PPQ-S&T - Identification Technology Program (ITP). Funding was provided by Section 10201 of the 2008 Farm Bill.
The authors would like to thank USDA-APHIS-PPQ-S&T - ITP, USDA-APHIS-PPQ - National Identification Services (NIS), and Colorado State
University for supporting the development of LepIntercept. This work would not have been possible without the help of numerous individuals,
which we attempt to list on this page. The efforts of everyone listed here are greatly appreciated, and we apologize in advance if anyone
who contributed to this project was omitted.
Terrence Walters (S&T - ITP, Fort Collins) supervised the project, allowed access to ITP's Visionary Digital imaging system, and provided
logistical and technical support. Joe Cavey (USDA-APHIS-PPQ - National Identification Services) provided resources needed to maintain and grow the PPQ Lepidoptera
insect and book collections that directly resulted in completion of this project. Facilities at the USFS in Delaware, Ohio for S. Passoa were
provided by Jim Slavicek. Facilities at USDA-S&T in Fort Collins, Colorado for T. Gilligan were provided by Richard Zink.
James Young provided extensive comments on fact sheets, keys, and consulted on numerous issues related to larval identification and
identification records. He literally made sure nothing fell through the cracks and was a champion every time there was a chance to do
something that benefited identifiers or this project.
The following persons and institutions provided loans or donations of larvae for morphological study:
John Brown, Michael Pogue, and Alma Solis, USDA-ARS - Systematic Entomology Laboratory, Smithsonian, Washington D.C.;
Estuardo Catalan, CENGICANA (Guatemalan Sugarcane Research and Training Center), Escuintla, Guatemala;
Marianne Horak, Australian National Insect Collection, CSIRO, Canberra;
Norman Johnson and Luciana Musetti, Charles A. Triplehorn Insect Collection, The Ohio State University;
Ricardo Antonio Polanczyk, Professor Assistente Doutor em Entomologia, UNESP Jaboticabal, Brazil;
Marja van der Straten, Plant Protection Service, Wageningen, Netherlands;
May Berenbaum and Terry Harrison, University of Illinois;
David Wagner, University of Connecticut;
Thomas Giles, USDA-PPQ, Nogales;
B. Christian Schmidt, Canadian National Collection of Insects, Ottawa.
Ella Gilligan (Loveland, Colorado) provided assistance with initial web page construction and tab functionality.
Michael Pogue and the following USDA-APHIS-PPQ Entomologists/Identifiers provided very helpful reviews of a beta version of this website and associated keys:
Lenis Fernando, Miami, Florida; Allan Smith-Pardo, S. San Francisco, California; James Young, Baltimore, Maryland; and Stephen Young, Blaine, Washington.
Alma Solis, Michael Pogue, and John Brown (USDA-ARS-SEL) allowed access to their unpublished identification aids, keys, manuscripts, and provided taxonomic support
on pyraloids, noctuoids, and tortricids, respectively.
Equipment provided by Joel Aronoff and Sue Ellis (USDA - Remote Pest Identification Program) was helpful in examining specimens used in this study.
Peter Touhey (USDA - NIS) provided access to interception records in PestID. Amanda Redford (S&T - ITP) assisted with technical support for the Lucid Key.
Jim Hayden (FSCA, McGuire Center) copied literature at a last minute request and assisted with pyraloid taxonomy.
George Godfrey's (Athens, Illinois) gift of his noctuid reprint collection greatly helped the reference section of this work. Equally important
was Charles Triplehorn's (The Ohio State University) reprint donations from his collection. Sadly, a few of S. Passoa's undergraduate and graduate
advisors who deserve thanks are not here to see this product. The late George Eickwort (Cornell University), Ellis MacLeod (University of Illinois),
and Dale Habeck (University of Florida) allowed access to larval collections at their institutions. This work would have been impossible without
notes taken at these collections or knowledge gained from them on morphology.
The following USDA-APHIS-PPQ Identifiers answered questions and contributed images or job aids to study and/or reference specimens that assisted greatly
with this project: G. Bartman, E. Bess, J. Botz, J. Brambila, C. Brodel, J. Brusch, N. Cottrell, T. Dobbs, R. Dones, J. Dooley, J Dorshorst, R. Elliott, G. Evans,
C. Gaona, P. Haslem, S.-H. Hung, R. Ito, M. Kharboutli, J. Korecki, D. Lee, L. Fernando, P. Johnson, B. Lindsey, S. Marmura, P. Marquez, N. Matteson, D. McCoy,
E. McDonald, K. Metz, N. Nieves, C. O'Donnell, C. Olsen, J. Orr, R. Pingel, H. Rasmussen, S. Romero, L. Saez, M. Segall, E. Serrano, T. Skarlinsky,
A. Smith-Pardo, W. Tang, R. Tracy, T. Watanabe, J. Weaver, E. White, W. Winnie, J. Young, S. Young, J. Zablotny, and J. Zhang.
Special thanks to M. Cazier Mosley, M. Kharboutli, H. Nadel, V. Mastro, and D. Lance for support of the Diatraea rearing project at the Otis Methods lab.
When the Weisman key update was submitted to a Lepidoptera-oriented journal in the 1990's by S. Passoa, it was refused with the reasoning that "if APHIS
won't publish it, that is a strike against the quality of the work." If that was a strike, then it seems that Terrence Walters and the Identification Technology
Program have hit a home run over the wall and out of the park with this project. We appreciate and never take this support for granted.
Materials and Methods
LepIntercept was produced as an identification resource for Lepidoptera larvae intercepted at U.S. ports of entry. Materials and methods used in the construction
of LepIntercept are listed in this section. For information how to use the resources found on this site, please click on the "How to Use LepIntercept" tab at the top of this page.
Taxa were selected for inclusion in the project by consulting the USDA's PestID database. The top 34 most commonly intercepted Lepidoptera taxa from the past ten years
(2002-2012) were selected as these make up more than 80% of the total number of larval interceptions during that time period. Spodoptera littoralis was added
to complete coverage of Spodoptera, bringing the total number of taxa to 35. A complete list of the taxa treated in LepIntercept is provided on the
"About LepIntercept" tab at the top of this page.
Information provided in the fact sheets is a combination of information pulled from the literature and personal observations. We follow the latest (2013) taxonomy for
all species, and, in some cases, this taxonomy is more recent than that used in USDA publications and databases. In these cases we provide background on the taxonomic
changes and cite or provide links to the appropriate literature. Tortricid taxonomy is based on the latest world catalogue at www.tortricid.net; pyraloid taxonomy is
based on the world database at www.pyraloidea.org; and noctuoid taxonomy is based on Lafontaine and Schmidt (2010), Zahiri et al. (2011), and other publications listed
directly on the individual fact sheets. Diagnostic characters listed for larvae are based on those listed in the literature and/or examination of
preserved specimens (see material examined). In the detailed larval diagnosis we attempt to review all relevant literature for a particular taxon group and also provide a summary
of our own findings. Host/origin information is a summary of the records listed in the USDA PestID database for a particular taxon. In a few cases, trade-sensitive records
have been removed. Country/location and host plant names are displayed as listed in the PestID database and have not been checked for accuracy, although obvious misspellings and
duplicates have been removed. The recorded distribution for each taxon was obtained from the literature in most instances. We use the term "New World" to refer to North America,
Central America, South America, and the Caribbean (not including Australasia). We do not include Mexico when referring to Central America. Identification authority is provided
as detailed information on the level of identification (family, genus, species) that should be attempted for interceptions of particular taxa from a particular zoogeographic
region. Pest characterization is based on three factors (taxonomy, distribution, and potential impact) outlined by Cavey (2001) that determines if a particular taxon is quarantine
significant for the U.S.
Our treatment of anomalous or doubtful host and origin records requires comment. No doubt some of these are true associations. An example is the confirmation of the
codling moth on citrus using molecular diagnostics. In other cases, it is hard to imagine the larva actually feeds on the intercepted host. There could be a misidentification
of either the host or the insect. Another possibility is that the larva may have been knocked off the true host in transit or the larva left the host while sitting on a
table at the port awaiting inspection. Steward  suggested not accepting extremely doubtful records (called conjectural data) in the PestID database and we generally
agree with his policy. We call attention to those records needing confirmation and omit the less likely data.
We examined more than 200 larval specimens, most of which were preserved in ethanol. A partial list of material examined is provided here. The following collection
abbreviations are used: USNM (United States National Museum/Smithsonian Institution collection, Washington, D.C.); SPIC (Steven C. Passoa collection, Columbus, Ohio);
TMG (Todd M. Gilligan collection, Loveland, Colorado); MVDS (Marja van der Straten collection, Wageningen, Netherlands).
In order to examine minute structures on larvae, at least one individual from nearly every taxon was dissected and slide mounted. A slit was cut in the larva from the anus to the
head capsule on the dorsal surface between the V setae using small Vannas spring scissors (Fine Science Tools; 2mm cutting edge). The larva was macerated in 10% KOH for
approximately one hour at 60 degrees Celsius and then transferred to a petri dish filled with distilled water. Mandibles were carefully removed using small Vannas scissors and forceps,
and the hypopharyngeal complex was removed by cutting a vertical slit next to each premental arm and a single horizontal slit below the prementum. The head was removed by cutting
around the base of the head capsule. The larval skin was cleaned using small brushes and forceps, and all structures were slide mounted in Euparal. More information on
preserving, staining, and examining larvae is found on the Larval Morphology page under the "Preserving and Studying Larvae" tab.
All photographs and figures were produced by the authors of LepIntercept (unless stated otherwise). In nearly all cases, preserved larvae were photographed
because these represent the type of larvae most often encountered by identifiers, and live larvae may appear very different. Macro photographs were taken using a
BK Lab or Passport Imaging System provided by the USDA-APHIS-PPQ-S&T - Identification Technology Program (ITP). Macro photos (of any non-slide mounted
subject) were produced by suspending the larva in a drop of "Astroglide personal lubricant" in a petri dish, and covering the larva with 70% ethanol. For several
close-up images, the larva was pinned into the bottom of a petri dish with a foam bottom and covered with 70% ethanol. Slide photographs were taken using
a Nikon DXM1200 microscope camera mounted on a Nikon Labophot microscope (or various other setups including a Nikon D90 mounted to a Martin Microscope MM
adaptor on a Leitz Aristoplan or Diaplan, Nikon 80i, or Zeiss Axiomat). Nearly all figures are a stack of 3-100 individual photos
produced using Zerene Stacker Professional 1.0. In Zerene Stacker, the
"PMax" method was used to stack most non-slide mounted subjects, and the "DMap" method was used to stack most slide mounted subjects. All figures were edited
using Adobe Photoshop CS5/6 Extended and scale bars were inserted using the Photoshop Analysis tools.
Illustrations and setal maps
All illustrations for the interactive identification key and larval setal maps were produced by the authors of LepIntercept using Adobe Illustrator CS5/6.
For character states figured in the identification key, actual photographs of structures were traced and/or simplified "overview" drawings of some structures were
produced to illustrate a particular character. For larval setal maps, a larva was dissected (see above), the skin was slide mounted, and a photograph of the skin
was imported into Adobe Illustrator to use as a template. Pinacula size, setal length, and relative setal position were inferred from the slide mounted skin and/or
detailed photographs of intact larvae.
The setal maps are designed to show the approximate position and number of "tactile primary setae" (see Stehr 1987: 302) for all species. We did not study the
thoracic leg or anal proleg chaetotaxy or any proprioceptors (MD, MSV, MV setae). For the primary setae, there are some differences in the setal length and relative
setal position compared to the published literature. There are several possible reasons for this variation. In most cases, the drawings were produced from a single
slide-mounted specimen and it is well known setal patterns can vary from individual to individual within a species or even differ on one side of the same specimen.
For example, the SV group on A1 of T. leucotreta is bisetose on several of our specimens, whereas it was illustrated as trisetose by Komai (1999) and Timm et al.
(2007). In addition, we did not examine the vouchers used to prepare any of the illustrations in published literature; there could be inaccuracies from the author,
or even misidentifications.
Larval setae are named according to the general setal map in Stehr (1987). Naming setal groups is generally clear, but trying to decide names within setal groups
(L1 or L2, SV1 or SV2, etc.) can be subject to reinterpretation among different families. This is especially true of the anal shield.
Slide preparations of larval skins have the advantage that they can be studied under a compound microscope so that small setae are less likely to be overlooked.
However, even with a slide-mounted skin, determining the presence or position of SD2 setae on the abdominal segments is often problematic. MacKay (1959) did
not show SD2 on A1-8 for either C. pomonella or G. molesta, whereas that seta is clearly present and easily visible in our specimens. We could not decide the
condition of SD2 in our preparation of T. ni - some abdominal segments apparently lacked SD2, but it was present on other parts of the body; in the end we
chose not to illustrate it. The inconspicuous SD2 in Pyrausta is present under high magnification but not shown on our map because it is basically "absent"
under low power.
A major disadvantage of using a slide-mounted skin is that some inaccuracies are inherent when trying to project a cylindrical larva on a flat surface.
MacKay (1959:11) stated that the positions of the D, SD, and L3 setae are especially hard to represent with this method. In many cases, slide mounting the setae
distorts their shape. As a result, we did not try to illustrate differences in setal thickness. For instance, the literature discusses SD1 being hairlike or
setaform on A9 in Gelechioidea and Noctuoidea (Stehr 1987) and there are other setae that need to be carefully examined. Noctuids may have the one
prothoracic L seta and SD1 on T2-3 thinner than normal. Diatraea appears to have SD1 on A9 hairlike, a character not yet studied in pyraloids.
Although these differences in setae may not be represented on the setal maps, they are mentioned in the fact sheet text if considered taxonomically
important. In all instances, we recommend consulting the habitus photographs in the fact sheets in addition to the setal maps to determine the position,
size, and shape of setae on intact specimens, remembering that setae can fall out of their socket. We do not recommend using the setal maps to "discover"
new characters to distinguish close relatives in difficult groups like Diatraea or Helicoverpa.
Copyright, Contact, Citation, and Disclaimers
LepIntercept was authored under a cooperative agreement between Colorado State University (CSU)
and the USDA-APHIS-PPQ-S&T - Identification Technology Program (ITP). This content may be freely distributed or copied in the public domain for
non-commercial purposes. However, it is requested that in any subsequent use of this work the authors, CSU, and ITP be given appropriate acknowledgement (see below).
LepIntercept may contain information, text, and images created and/or prepared by individuals or institutions other than CSU or USDA-APHIS that may be protected
by copyright. Sources of information and text are mentioned on the Acknowledgments tab, and in most instances the origin of images has been indicated on specific
fact sheets. Users must seek permission from the copyright owner(s) to use this material; contact the authors (above) if you need assistance identifying or locating
the copyright owner for a particular image or have any questions related to use of information on this site.
LepIntercept was authored by Dr. Todd M. Gilligan and Dr. Steven C. Passoa. Todd is a Research Scientist in the Bioagricultural Sciences and Pest
Management Department at Colorado State University in Fort Collins, CO. Steven is the National Lepidoptera Specialist for USDA-APHIS-PPQ and is located at
The Ohio State University in Columbus, OH. Unless otherwise noted, all web pages (including design and graphics), photographs, illustrations, keys, and other
documents for LepIntercept were produced by the authors. If you have any questions regarding LepIntercept, please contact
Todd Gilligan [click here] for information on Tortricidae, web page content, images, or any general questions about
this site. Please contact Steven Passoa [click here] for taxonomic questions on other groups,
literature citations, and information on APHIS policies (identification authority, etc.).
Gilligan, T. M. & S. C. Passoa. 2014. LepIntercept, An identification resource for intercepted Lepidoptera larvae.
Identification Technology Program (ITP), USDA-APHIS-PPQ-S&T, Fort Collins, CO. [accessed at www.lepintercept.org].
Photo: T. M. Gilligan & S. C. Passoa, LepIntercept (www.lepintercept.org)
(Unless otherwise noted)
While the authors have made every effort to provide accurate information in LepIntercept, the authors, Colorado State University,
and USDA-APHIS-PPQ-S&T specifically disclaim all legal liability with respect to the accuracy, completeness,
or usefulness of any information contained in LepIntercept. The authors and associated institutions shall assume no legal liability for any damages,
including direct, indirect, consequential, compensatory, special, punitive, or incidental damages arising from or relating to the use of LepIntercept
or the information and materials provided by or linked from LepIntercept.
Some web pages in LepIntercept provide links to Internet sites for the convenience of users. The authors, CSU, and USDA-APHIS-PPQ-S&T are not
responsible for the availability or content of these external sites, nor do the authors, CSU, and USDA-APHIS-PPQ-S&T endorse or warrant the products,
services, or information described or offered by these Internet sites.