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Aguiar, C.M., C.A. Garafálo, and G.F. Almeida. 2005. Trap-nesting bees (Hymenoptera: Apoidea) in areas of dry semideciduous forest and caatinga, Bahia, Brazil. Revista Brasileira de Zoologia 22: 1030–1038.

Ahmad F. & P. Roy. 2000. Bhutan: Indigenous Honey Bee Project Study- travel reports. ICIMOD organization, Kathmandu. Available online at: http://www.icimod.org/?q=1511

Alcock J., G.C. Eickwort, and K.R. Eickwort. 1977. The reproductive behavior of Anthidium maculosum (Hymenoptera: Megachilidae) and the evolutionary significance of multiple copulations by females. Behavioral Ecology and Sociobiology 2: 385–396.

Alfken, J.D. 1914 Beitrag zur Kenntnis der Bienenfauna von Algerien. Mémoires de la Société Entomologique Belgique 22: 185–237.

Allen, M.F. 1995. Bees and beekeeping in Nepal. Bee World 76: 185–194.

Alvarez, L.J., M. Lucia, S.P. Durante, L.M. Passarelli, and V.H. Gonzalez. 2015. Description of the male, new geographical record, and biological notes on the small resin bee Hypanthidioides (Ctenanthidium) bifasciata (Urban, 1993) (Hymenoptera: Megachilidae). The Pan-Pacific Entomologist 91: 184–188.

Alves-dos-Santos, I. 2003. Trap-nesting bees and wasps on the University Campus in São Paulo, Southeastern Brazil (Hymenoptera, Aculeata). Journal of the Kansas Entomological Society 76: 328–334.

Alves-dos-Santos, I. 2004. Biologia de nidificação de icação de Anthodioctes megachiloides Holmberg (Anthidiini, Megachilidae, Apoidea). Revista Brasileira de Zoologia 21: 739–744.

Alves-dos-Santos, I. 2010. Anthodioctes megachiloides nesting in an abandoned nest of Brachymenes dyscherus (Hymenoptera: Megachilidae; Vespidae). Journal of the Kansas Entomological Society 83: 39–91.

Amiet, F., M. Herrmann, A. Mueller, and R. Neumeyer. 2004. Apidae 4: Anthidium, Chelostoma, Coelioxys, Dioxys, Heriades, Lithurgus, Megachile, Osmia, Stelis. Fauna Helvetica 9: 1–273.

Anderson, D.L. and M.J. Morgan. 2007. Genetic and morphological variation of bee-parasitic Tropilaelaps mites (Acari: Laelapidae). Experimental and Applied Acarology 43: 1–43.

Arias M. C. and W. S. Sheppard. 1996. Molecular phylogenetics of honeybee subspecies (Apis mellifera L.) inferred from mitochondrial DNA sequences. Molecular Phylogenetics and Evolution 5: 557–566.

Arias, M.C. and W.S. Sheppard. 2005. Phylogenetic relationships of honey bees (Hymenoptera: Apinae: Apini) inferred from nuclear and mitochondrial DNA sequence data. Molecular Phylogenetics and Evolution 37: 25–35.

Arias, M.C., S. Tingek, A. Kelitu, and W.S. Sheppard, 1996. Apis nuluensis Tingek, Koeniger, and Koeniger, 1996 and its genetic relationship with sympatric species inferred from DNA sequences. Apidologie 27: 415–422.

Armbruster, W. S. and  D. A. Guinn. 1989. The solitary bee fauna (Hymenoptera: Apoidea) of interior and arctic Alaska: flower associations, habitat use, and phenology. Journal of the Kansas Entomological Society, 468-483.

Armbruster, S.W. 1984. The role of resin in angiosperm pollination: ecological and chemical considerations. American Journal of Botany 71: 1149–1160.

Armbruster, S.W. and G.L. Webster. 1981. Sistemas de polinizavao de duas especies simpatricas de Dalechampia (Euphorbiaceae) no Amazonas, Brasil. Acta Amazonica 11: 13–17.

Ascher, J.S. and J. Pickering. 2016. Discover Life bee species guide and world checklist (Hymenoptera: Apoidea: Anthophila). Discover Life. https://www.discoverlife.org/mp/20q?search=Apoidea

Ascher, J.S. and J. Pickering. 2016. Discover Life bee species guide and world checklist: Bathanthidium. Discover Life. https://www.discoverlife.org/mp/20q?search=Bathanthidium

Augusto, S.C. and C.A. Garófalo. 1998. Behavioral aspects of Hoplostelis bilineolata (Spinola) (Hymenoptera, Megachilidae), a cleptoparasite of Euglossa cordata (Linnaeus) (Hymenoptera, Apidae), and the behavior of hosts in parasitized nests. Revista Brasileira de Entomologia 41: 507–515.


Baker, J.R. 1971. Development and sexual dimorphism of larvae of the bee genus Coelioxys. Journal of the Kansas Entomological Society 44: 225–235.

Baker, D.B. 1995. A review of the Asian species of the genus Euaspis Gerstäcker (Hymenoptera: Apoidea: Megachilidae). Zoologische Mededelingen, 69: 281–302.

Baker, D.B. 1996. An annotated list of the nominal species assigned to the genus Afrostelis Cockerell (Hymenoptera, Apoidea, Megachilidae). Deutsche Entomologische Zeitschrift 43: 155–157.

Baker, D.B. 1999. On new stelidine bees from S.W. Asia and N.W. Africa, with a list of the Old-World taxa assigned to the genus Stelis Panzer, 1806 (Hymenoptera, Apoidea, Megachilidae). Deutsche Entomologische Zeitschrift, 46: 231–242.

Baker, D.B. and Engel, M.S. 2006. A new subgenus of Megachile from Borneo with arolia (Hymenoptera: Megachilidae). American Museum Novitates 3505: 1–12.

Banaszak, J. and L. Romasenko. 1998. Megachilid bees of Europe (Hymenoptera, Apoidea, Megachilidae). Bydgoszcz University, Bydgoszcz, Poland.

Banaszak J. and L. Romasenko. 2001. Megachilid bees of Europe. Second edition. Bydgoszcz: Bydgoszcz University Press. 239 pp.

Barthell, J.F., G.W. Frankie, and R.W. Thorp. 1998. Invader effects in a community of cavity nesting megachilid bees (Hymenoptera: Megachilidae). Environmental Entomology 27: 240–247.

Basavarajappa S., and K.S. Raghunandan. 2013. Colony status of Asian giant honeybee, Apis dorsata Fabricius in Southern Karnataka, India. African Journal of Agricultural Research 8 (8): 680–689.

Batra, S.W.T. 1976. Comparative efficiency of alfalfa pollination by Nomia melanderi, Megachile rotundata, Anthidium florentinum, and Pithitis smaragdula (Hymenoptera: Apoidea). Journal of the Kansas Entomological Society 49: 18–22.

Batra, S.W.T. 1978. Aggression, territoriality, mating, and nest aggregation of some solitary bees (Hymenoptera: Halictidae, Megachilidae, Colletidae, Anthophoridae). Journal of the Kansas Entomological Society 51: 547–559.

Batra, S.W.T. 1979. Osmia cornifrons and Pithitis smaragdula, two Asian bees introduced into the United States for crop pollination. Food and Agricultural Organization of the United Nations.

Batra S. 1995. Biology of Apis laboriosa Smith, a pollinator of apples at high altitudes in the greater Himalaya of Gahrwal, India. Tektran. United States Department of Agriculture, Agricultural Research Service.

Baumann, J.M., K. Walker, C. Threlfall, and N.S. Williams. 2016. African carder bee, Afranthidium (Immanthidium) repetitum Hymenoptera: Megachilidae): A new exotic species for Victoria. The Victorian Naturalist 133 (1), p.21.

Bennett, F.D. 1966. Notes on the biology of Stelis (Odontostelis) bilineolata (Spinola), a parasite of Euglossa cordata (Hymenoptera: Apidae: Euglossini). Journal of the New York Entomological Society 74: 72–79.

Bezabih G., N. Adgaba, H.R. Hepburn, and C.W.W. Pirk. 2014. The territorial invasion of Apis florea in Africa. African Entomology 22 (4): 888–890.

Bosch, J., W.P. Kemp, and S.S. Peterson. 2000. Management of Osmia lignaria (Hymenoptera: Megachilidae) populations for almond pollination: methods to advance bee emergence. Environmental Entomology 29: 874–883.

Bosch, J., W.P. Kemp, and G.E. Trostle. 2006. Bee population returns and cherry yields in an orchard pollinated with Osmia lignaria (Hymenoptera: Megachilidae). Journal of Economic Entomology 99: 408–413.

Bosch, J., N. Vicens, and M. Blas. 1993. Analisis de los nidos de algunos Megachilidae nidificantes en cavidades pre-establecidas. Orsis 8: 53–63.

Bosch, J., F. Sgolastra, and W.P. Kemp. 2008. Life cycle ecophysiology of Osmia mason bees used as crop pollinators. Bee Pollination in Agricultural Ecosystems. Oxford University Press, New York, pp. 83–104.

Buck, M., S.M. Paiero, and S.A. Marshall. 2005. New records of native and introduced aculeate Hymenoptera from Ontario, with keys to eastern Canadian species of Cerceris (Crabronidae) and eastern Nearctic species of Chelostoma (Megachilidae). Journal of the Entomological Society of Ontario 136: 37–52.


Camarotti-de-Lima, M. and C.F. Martins. 2005. Biologia de nidificação e aspectos ecológicos de Anthodioctes lunatus (Smith) (Hymenoptera: Megachilidae, Anthidiini) em área de tabuleiro nordestino, PB. Neotropical Entomology 34: 375–380.

Cane, J.H. 1996. Nesting resins obtained from Larrea pollen host by an oligolectic bee, Trachusa larreae (Cockerell). Journal of the Kansas Entomological Society 69: 99–102.

Cane, J.H., T. Griswold, and F.D. Parker. 2007. Substrates and materials used for nesting by North American Osmia bees (Hymenoptera: Apiformes: Megachilidae). Annals of the Entomological Society of America 100: 350–358.

Cane, J.H. 2016. Specialist bees collect Asteraceae pollen by distinctive abdominal drumming (Osmia) or tapping (Melissodes, Svastra). Arthropod-Plant Interactions 11: 257–261.

Chaiyawong T., S. Jayasvasti, G. Suwannapong, and S. Wongsiri. 2001. Scanning electron microscopic analysis of honey bees, Apis nigrocincta, A. cerana, A. koschevnikovi and A. mellifera stings. In Wongsiri, S. (ed.), Proceedings of the Seventh International Conference on Tropical Bees: Management and Diversity & Fifth Asian Apicultural Association Conference 19–25 March, Thailand. pp. 121–128.

Chaiyawong T., S. Deowanish, S. Wongsiri, H.E. Sylvester, T.E. Rinderer, and L. de Guzman. 2004. Multivariate morphometric study of Apis florea in Thailand. Journal of Apicultural Research 43 (3): 123–127.

Cilla, G. and G. Rolón. 2012. Macroscopic and microscopic studies of the nests and the stages involved in the nesting process of Centris muralis Burmeister (Hymenoptera: Apidae: Centridini) bee in the adobe walls, in La Rioja, Argentina. Biologia 67: 573–583.

Claude-Joseph, F. 1926. Recherches biologiques sur les Hyménoptères du Chili (Mellifères). In Annales des Sciences Naturelles, Zoologie 9 (10): 113–268.

Cockerell, T.D.A. 1909. Some new bees and other notes. The Canadian Entomologist 41: 128–131.

Cocom Pech, M.E., W.J. May-Itzá, L.A. Medina, and J.J.G. Quezada-Euán. 2008. Sociality in Euglossa (Euglossa) viridissima Friese (Hymenoptera, Apidae, Euglossini). Insectes Sociaux 55: 428–433.

Cordeiro, G.D., A.L.O. Nascimento, I. Alves-dos-Santos, and C.A. Garófalo. 2015. The cleptoparasitic bee Austrostelis iheringi (Schrottky, 1910) (Anthidiini, Megachilidae): new geographical records and potential hosts. Journal of the Kansas Entomological Society 88: 453–456.

Custer, C.P. and C.H. Hicks. 1927. Nesting habits of some anthidiine bees. Biological Bulletin 52: 258–277.

Crosswhite, F.S. and C.D. Crosswhite. 1966. Insect pollinators of Penstemon series Graciles (Scrophulariaceae) with notes on Osmia and other Megachilidae. American Midland Naturalist 76: 450–467.


Damus M.S. and G.W. Otis. 1997. A morphometric analysis of Apis cerana F. and Apis nigrocincta Smith populations from Southeast Asia. Apidologie 28 (5): 309–323.

Davidson, A. 1895. The habits of Californian bees and wasps. Anthidium emarginatum, its life history and parasites. Entomological News 6: 252–253.

Dieringer, G., T.P. Ramamoorthy, and P.T. Lezama. 1991. Floral visitors and their behavior to sympatric Saliva species (Lamiaceae) in Mexico. Acta Botanica Mexicana 13: 75–83.

Discover Life. 2018. Anthidium (Gulanthidium). Discover Life. https://www.discoverlife.org/mp/20p?see=Gulanthidium&name=Anthidium&flags=subgenus:&mobile=1

Discover Life. 2018b. Anthidium oblongatum (Illiger, 1806). Discover Life. https://www.discoverlife.org/20/q?search=Anthidium+oblongatum

Discover Life. 2018c. Anthidium florentinum (Fabricus, 1775). Discover Life. https://www.discoverlife.org/mp/20q?search=Anthidium+florentinum&flags=subgenus

Doroshina, L.P. 1990. Protection from chalcids and optimization of nesting conditions for megachilid bees (Hymenoptera, Apoidea, Megachilidae). Zoologicheskii Zhurnal 69: 55–59.

Droege, S. 2015. The Very Handy Manual: How to Catch and Identify Bees and Manage a Collection. USGS Native Bee Inventory and Monitoring Lab (https://www.pwrc.usgs.gov/nativebees/Handy%20Bee%20Manual/The%20Very%20Handy%20Manual%20-%202015.pdf).

Droege, S.W. and L.H. Shapiro. 2011. An August survey of wild bees (Hymenoptera: Apoidea) in the northeastern port areas of Baltimore, Maryland and the second North American record of Pseudoanthidium nanum (Mocsáry). The Maryland Entomologist 5: 33–44.

Ducke, A. 1900. Die Bienengattung Osmia Panz. Ber. naturw.-med. Ver. Innsbruck 25: 1–323.

Dumesh and Packer. 2013. Three new species of Neofidelia (Hymenoptera: Apoidea: Megachilidae) from Northern Chile. Zootaxa 3609 (5): 471–483.

Dung P.X., P.H. Thai, and L.Q. Trung. 1996. Some comparative characteristics between Apis florea and Apis andreniformis in Minh Hai Province, Proceedings of the 3rd Asian Apiculture Association, Hanoi. pg. 62.


Eardley, C. 2013. A taxonomic revision of the southern African leaf-cutter bees, Megachile Latreille sensu stricto and Heriadopsis Cockerell (Hymenoptera: Apoidea: Megachilidae). Zootaxa 3601 (1): 1–133.

Eardley, C. and T. Griswold. 2016. Taxonomic revision of the Afrotropical bee genus Serapista Cockerell (Hymenoptera: Apoidea: Megachilidae: Megachilinae: Anthidiini). Zootaxa 4111: 334–364.

Eickwort, G.C. 1970. Hoplitis anthocopoides, a European mason bee established in New York state (Hymenoptera: Megachilidae).

Engel, M.S. 1999. The taxonomy of recent and fossil honey bees (Hymenoptera: Apidae; Apis). Journal of Hymenoptera Research 8: 165–196.

Engel, M.S. 2002. Phylogeny of the bee tribe Fideliini (Hymenoptera: Megachilidae), with the description of a new genus. African Entomology 10: 305–313.

Engel, M.S. 2004. Fideliine phylogeny and classification revisited (Hymenoptera: Megachilidae). Journal of the Kansas Entomological Society 77: 821–836.

Engel M. S. 2012. The honey bees of Indonesia (Hymenoptera: Apidae). Treubia 39: 41–49.

Erbar, C. and P. Leins. 2017. Sex and breeding behavior of the Sicilian snail-shell bee (Rhodanthidium siculum Spinola, 1838; Apoidea-Megachilidae): preliminary results. Arthropod-Plant Interactions 11: 317–328.

Evans, H.E. 1993. Observations on the nests of Paranthidium jugatorium perpictum (Cockerell) (Hymenoptera: Megachilidae: Anthidiini). The Pan-Pacific Entomologist 69: 319–322.


Fabricius, J.C. 1793. Entomologia Systematica. Hafniae, Proft. Vol. II, 8+519pp.

Falamarzi, S., B. Habibpour, M.S. Mossadegh, and A. Monfared. 2017. Species inventory of Megachilidae (Hymenoptera: Apoidea) in south of Fars province, Iran. Entomofauna 38: 89–104.

Food and Agriculture Organization of the United Nations, Rome (FAOUNR). 2008. Tools for conservation and use of pollination services. Pollination Services for Sustainable Agriculture: 1–143.

Ferguson, W.E. 1962. Biological characteristics of the mutillid subgenus Photopsis Blake and their systematic values (Hymenoptera). University of California Publications in Entomology 27: 1–92.

Forrest, J.R.K. and S.P.M. Chisholm. 2016. Direct benefits and indirect costs of warm temperatures for high-elevation populations of a solitary bee. Ecology 98: 359–369.

Frank P., L. Garnery, A. Loiseau, B.P. Oldroyd, H.R. Hepburn, M. Solignac et al. 2001. Genetic diversity of the honeybee in Africa: microsatellite and mitochondrial data. Heredity 86: 420–430.

Friese, H. 1923. Die europäischen Bienen (Apidae). Das Leben und Wirken unserer Blumenwespen. Berlin: Walter de Gruyter. 456 pp.

Frohlich, D.R. 1983. On the nesting biology of Osmia (Chenosmia) bruneri. Journal of the Kansas Entomological Society 56: 123–130.

Fuchs S., N. Koeniger, and S. Tingek. 1996. The morphometric position of A. nuluensis (Tingek, Koeniger, and Koeniger) within cavity-nesting honey bees. Apidologie 27: 397–406.


Gan Y.Y., G.W. Otis, M.B. Mardan, and S.G. Tan. 1991. Allozyme diversity in Asian Apis, in: Smith D.R. (Ed.), Diversity in the genus Apis. Westview Press, Boulder, Co. pp.: 117–130.

Garnery L., J.M. Cornuet, and M. Solignac. 1992. Evolutionary history of the honeybee Apis mellifera inferred from mitochondrial DNA analysis. Molecular Ecology 1: 145–154.

Garófalo, C.A., E. Camillo, J.O. Campos, and J.C. Serrano. 1992. Nest re-use and communal nesting in Microthurge corumbae (Hymenoptera: Megachilidae), with special reference to defense. Insectes Sociaux 39: 310–311.

GBIF.org. 14 June 2019. GBIF Occurrence Download https://doi.org/10.15468/dl.wmukot

GBIF.org. 11 July 2019. GBIF Occurrence Download. https://doi.org/10.15468/dl.lw8pwz

GBIF.org. 8 July 2019. GBIF Occurrence Download. https://doi.org/10.15468/dl.ytygqt

GBIF.org. 12 July 2019. GBIF Occurrence Download. https://doi.org/10.15468/dl.2gbgt2

GBIF.org. 8 July 2019. GBIF Occurrence Download. https://doi.org/10.15468/dl.bfuama

GBIF.org. 19 June 2019. GBIF Occurrence Download. https://doi.org/10.15468/dl.xi3aul

GBIF.org . 12 July 2019. GBIF Occurrence Download. https://doi.org/10.15468/dl.g5ar8l

GBIF.org. 26 July 2019. GBIF Occurrence Download. https://doi.org/10.15468/dl.itllyg

Gess, S.K. and F.W. Gess. 2007. Notes on nesting and flower visiting of some anthidiine bees (Hymenoptera: Megachilidae: Megachilinae: Anthidiine) in southern Africa. Journal of Hymenoptera Research 16: 30–50.

Gess, S.K. and F.W. Gess. 2008. Patterns of usage of snail shells for nesting by wasps (Vespidae: Masarinae and Eumeninae) and bees (Megachilidae: Megachilinae) in Southern Africa. Journal of Hymenoptera Research 17: 86–109.

Gess, S.K., F.W. Gess, and R.W. Gess. 2008. Patterns of usage of snail shells for nesting by wasps (Vespidae: Masarinae and Eumeninae) and bees (Megachilidae: Megachilinae) in Southern Africa. Journal of Hymenoptera Research 17: 86–109.

Gess, S.K. and P.A. Roosenschoon. 2017. Notes on the nesting of three species of Megachilinae in the Dubai Desert Conservation Reserve, UAE. Journal of Hymenoptera Research 54: 43–56.

Giles, V. and J.S. Ascher. 2006. A survey of the bees of the Black Rock Forest Preserve, New York (Hymenoptera: Apoidea). Journal of Hymenoptera Research 15: 208–231.

Gogala, A., B, Surina. 2011. Foraging behaviour of the bee Osmia apicata Smith, 1853 (Hymenoptera: Megachilidae). Acta entomologica Slovenica 19: 139–144.

Gonzalez, V.H. 2008. Phylogeny and classification of the bee tribe Megachilini (Hymenoptera: Apoidea, Megachilidae), with emphasis on the genus Megachile. Doctoral dissertation, University of Kansas.

Gonzalez, V.H. 2013. Taxonomic comments on Megachile subgenus Chrysosarus (Hymenoptera: Megachilidae). Journal of Melittology 5: 1–6.

Gonzalez, V.H. and T.L. Griswold. 2011. Taxonomic notes on the small resin bees Hypanthidioides subgenus Michanthidium (Hymenoptera, Megachilidae). ZooKeys 177: 5–58.

Gonzalez, V.H. and T.L. Griswold. 2013. Wool carder bees of the genus Anthidium in the Western Hemisphere (Hymenoptera: Megachilidae): diversity, host plant associations, phylogeny, and biogeography. Zoological Journal 168: 221–425.

Gonzalez, V.H., M.S. Engel, and T. Griswold. 2013. The lithurgine bees of Australia (Hymenoptera: Megachilidae), with a note on Megachile rotundipennis. Journal of Melittology 11: 1–19.

Gonzalez, V.H., T. Griswold, C.J. Praz, B.N. Danforth. 2012. Phylogeny of the bee family Megachilidae (Hymenoptera: Apoidea) based on adult morphology. Systematic Entomology 37: 261–286.

Gorain, M., S.K. Charan, and S.I. Ahmed. 2012. Role of insect bees in the pollination of Prosopis cineraria (L.) Druce (Leguminosae, Subfamily Mimosoideae) in Rajasthan. Advances in Applied Science Research 3: 3448–3451.

Gotlieb, A., G. Pisanty, J.G. Rozen Jr., A. Müller, G. Röder, C. Sedivy, and C. Praz. 2014. Nests, floral preferences, and immatures of the bee Haetosmia vechti (Hymenoptera: Megachilidae: Osmiini). American Museum Novitates 3803: 1–20.

Grace, A. 2010. Introductory biogeography to bees of the eastern Mediterranean and Near East. Bexhill Museum, Sussex, United Kingdom.

Graeffe, E. 1902. Die Apiden-Fauna des österreichischen Küstenlandes. Verh. k. k. zool.-bot. Ges. Wien, 52: 113–135.

Graham, K.K. 2018. Heterospecific Hymenoptera found inside the nests of Bombus impatiens (Hymenoptera: Apidae). The Great Lakes Entomologist 51: 1–4.

Grigarick, A.A. and L.A. Stange. 1968. Pollen collecting bees of the Anthidiini of California (Hymenoptera: Megachilidae). Bulletin of the California Insect Survey 9: 1–113.

Griswold, T. 1985. A generic and subgeneric revision of the Heriades genus-group. Logan: Ph.D. thesis, Utah State University. xiii + 165 pp.

Griswold, T. 1986. Notes on the nesting biology of Protosmia (Chelostomopsis) rubifloris (Cockerell) (Hymenoptera: Megachilidae). The Pan-Pacific Entomologist 62: 84–87.

Griswold, T.L. 1986. A new heriadine bee from the Mojave Desert. The Southwestern Entomologist 11: 165–169.

Griswold, T.L. 1991. A review of the genus Microthurge (Hymenoptera: Megachilidae). Pan-Pacific Entomologist 67: 115–118.

Griswold, T.L. 1994. A review of the Ochreriades. Pan-pacific Entomologist 70: 318–321.

Griswold, T.L. 1994. Taxonomic notes on some heriadines, with descriptions of three new species (Hymenoptera: Megachilidae). Journal of the Kansas Entomological Society 67: 17–28.

Griswold, T. 2015. A review of Trachusoides Michener and Griswold (Hymenoptera: Megachilidae). Zootaxa 3949: 147–150.

Griswold, T.L. 2018. First record of Stenoheriades Cockerell in tropical Asia: Stenoheriades bifida, new species (Hymenoptera: Megachilidae). Zootaxa 4370: 279–282.

Griswold, T. and V.H. Gonzalez. 2011. New species of the Eastern Hemisphere genera Afroheriades and Noteriades (Hymenoptera, Megachilidae), with keys to species of the former. Zookeys 159: 65–80.

Griswold, T.L. and V.H. Gonzalez. 2013. A new species of the rare African wool-carder bee genus Anthidioma (Hymenoptera: Megachilidae) from Namibia. African Entomology 21: 117–180.

Griswold, T.L. and C.D. Michener. 1997. The classification of the Osmiini of the Eastern Hemisphere (Hymenoptera, Megachilidae). Journal of the Kansas Entomological Society 70: 207–253

Griswold, T. and F.D. Parker. 2003. Stelis rozeni, new species, the first record of the parasitic bee genus Stelis from Southern Africa (Hymenoptera: Megachilidae). Journal of the Kansas Entomological Society 76: 282–285.

Griswold, T. and M.G. Rightmyer. 2017. A revision of the subgenus Osmia (Diceratosmia), with descriptions of four new species (Hymenoptera: Megachilidae). Zootaxa 4337: 1–37.

Guisse, J.K and D.G. Miller. 2011. Distribution and habitat preferences of Osmia lignaria (Hymenoptera: Megachilidae) and associated cavity-nesting insects in California’s Sierra Nevada foothills adjacent to the Sacramento Valley. The Pan-Pacific Entomologist 87 (3): 188–195.

Gupta, R.K. 2014. Taxonomy and distribution of different honeybee species. In: Gupta R.K., Reybroeck W., van Veen J. W. & A. Gupta (eds): Beekeeping for poverty alleviation and livelihood security Vol.1: Technological aspects of beekeeping. Pp. 63–103.


Haddad, N. S. Fuchs, H.R. Hepburn, and S.E. Radloff. 2009. Apis florea in Jordan: source of the founder population. Apidologie 40: 508–512.

Hadisoesilo S. and G.W. Otis. 1996. Drone flight times confirm the species status of Apis nigrocincta Smith, 1861 to be a species distinct from Apis cerana F, 1793, in Sulawesi, Indonesia. Apidologie 27 (5) 361–369.

Hadisoesilo S. and G.W. Otis. 1998. Differences in drone cappings of Apis cerana and Apis nigrocincta. Journal of Apicultural Research 37: 11–15.

Hadisoesilo S., G.W. Otis, and M. Meixner M. 1995. Two distinct populations of cavity-nesting honeybees (Hymenoptera: Apidae) in South Sulawesi, Indonesia. Journal of the Kansas Entomological Society 68: 399–407.

Hadisoesilo S., G.W. Otis G.W., M. Meixner. 1998. Differences in drone cappings of Apis cerana and Apis nigrocincta. Journal of Apicultural Research 37: 11–15.

Hadisoesilo S., R. Raffiudin. W. Susanti, T. Atmowidi, C. Hepburn, S. Radloff, S. Fuchs, and R.H. Hepburn. 2008. Morphometric analysis and biogeography of Apis koschevnikovi Enderlein (1906). Apidologie 39: 495–503.

Haeseler, V. 2008. Zur Biologie der mediterranen Mauerbienen Osmia balearica Schmiedeknecht 1885 und Osmia uncicornis Pérez 1895 (Hymenoptera: Apidae). Abhandlungen des Naturwissenschaftlichen Vereins zu Bremen 46: 289–300.

Haider, M. S. Dorn, C. Sedivy, A. Müller. 2013. Phylogeny and floral hosts of a predominantly pollen generalist group of mason bees (Megachilidae: Osmiini). Biological Journal of the Linnean Society 111: 78–91.

Hepburn H.R. and C. Hepburn. 2007. Bibliography of Apis koschevnikovi Enderlein (1906). Apidologie 38 (6): 507.

Hepburn, H.R. and S.E. Randloff. 2011. Honeybees of Asia. Springer, Berlin. Xii+669p.

Hepburn, H.R., B. Radloff, G.W. Otis, S. Fuchs, L.R. Verma, T. Ken, T. Chaiyawong, G. Tahmaseb, R. Ebadi, and S. Wongsiri. 2005. Apis florea: morphometrics, classification and biogeography. Apidologie 36: 359–376.

Hicks, C.H. 1926. Nesting habits and parasites of certain bees of Boulder County, Colorado. University of Colorado Bulletin 15: 217–252.

Hicks, C.H. 1929. On the nesting habits of Callanthidium illustre (Cresson). The Canadian Entomologist 61: 1–8.

Hicks, C.H. 1928. Notes on Anthidium palliventre Cresson. The Pan-Pacific Entomologist 5: 51–52.

Hicks, B. 2009. Observations on the nest structure of Osmia inermis (Hymenoptera: Megachilidae) from Newfoundland, Canada. Journal of the Acadian Entomological Society 5: 12–18.

Hoebeke, E.R. and A.G. Wheeler, Jr. 1999. Anthidium oblongatum (Illiger): an old world bee (Hymenoptera: Megachilidae) new to North America, and new North America records for another adventive species, A. manicatum (L.). The University of Kansas Natural History Museum Special Publication 24: 21–24.

Horning, D.S. 1969. First recorded occurrence of the genus Callanthidium in Idaho with notes on three nests of C. formosum (Cresson) (Hymenoptera: Megachilidae). The Pan-Pacific Entomologist 45: 239.

Hurd, P.D, Jr. 1958. The American bees of the genus Dioxys Lepeletier and Serville (Hymenoptera: Megachilidae). University of California Publications in Entomology 14: 275–302

Hurd Jr., P.D. 1979. Superfamily Apoidea: In: Krombein K.V, J.P.D. Hurd, D.R. Smith, B.D. Burks, eds Catalog of Hymenoptera in America North of Mexico. Vol. 2. Washington, DC: Smithsonian Institution Press: 1741–2209.

Hurd, P.D. and C.D. Michener. 1955. Megachiline Bees of California (Hymenoptera: Megachilidae). Bulletin of the California Insect Survey (Vol. 3). Berkeley and Los Angeles: University of California Press, 247 pp.


Invasive Species Council of Australia. 2014. Case 12: biosecurity failures in Australia: Asian honey bee. Available at: https://invasives.org.au/wp-content/uploads/2014/09/Biosecurity-failures-asian-honey-bees.pdf.

ITIS. Retrieved [Jul, 12, 2017], from the Integrated Taxonomic Information System on-line database, http://www.itis.gov.

Ivanov, S.P. 2006. The nesting of Osmia rufa (L.) (Hymenoptera, Megachilidae) in the Crimea: Structure and composition of nests. Entomological Review 86: 524–533.

Ivanov, S.P., A.V. Fateryga, and M.A. Kobetskaya. 2013. The nesting biology of the bee, Osmia dimidiata Morawitz, 1870 (Hymenoptera, Megachilidae) in the Crimea. Entomological Review 93: 675–694.

Iwata, K. 1976. Evolution of Instinct: Comparative Ethology of Hymenoptera. Amerind Publishing Co. Pvt. Ltd., New Delhi, 536 pp.


Jaycox, E.R. 1966. Observations on Dioxys productus productus (Cresson) as a parasite of Anthidium utahense Swenk (Hymenoptera: Megachilidae). The Pan-Pacific Entomologist 42: 18–20.

Jaycox, E.R. 1967. Territorial behavior among males of Anthidium banningense (Hymenoptera: Megachilidae). Journal of the Kansas Entomological Society 40: 565–570.

Jayasvasti, S., and S. Wongsiri. 1992. Scanning electron microscopy analysis of honeybees (Apis florea, Apis dorsata, Apis cerana, Apis mellifera, Apis andreniformis and Apis koschevnikovi) stings. Recent Advances in Toxicology Research 2: 193–204.

Jayasvasti, S., and S. Wongsiri. 1993. Scanning electron microscopy analysis of honey bee - stings of six species (Apis florea, Apis dorsata, Apis cerana, Apis koschevnikovi, Apis florea, and Apis andreniformis). Honeybee Science 14: 105–109.

Johnson, S.A. 1904. Nests of Anthidium illustre Cress. Entomological News 15: 284.


Kandori, I., M. Tamaru, and T. Yokoi. 2010. Nesting habits of a pollinator: Osmia orientalis (Hymenoptera: Megachilidae). Japanese Journal of Applied Entomology and Zoology 54: 77–84.

Keeling C., G.W. Otis, S. Hadisoesilo, and K. Slessor. 2001. Mandibular gland component analysis in the head extracts of Apis cerana and Apis nigrocincta. Apidologie 32 (3): 243–252.

Keshtkar, A., A. Monfared, and M. Haghani. 2015. A survey on pollinator bees (Hymenoptera: Apoidea) in parks and gardens of Shiraz City, Iran. Entomofauna 36: 53–64.

Khodaparast, R. and A. Monfared. 2012. A survey of bees (Hymenoptera: Apoidea) from Fars Province, Iran. Zootaxa 3445: 37–58.

Khodaparast, R. and A. Monfared. 2013. Taxonomic studies on Osmiine bees (Hymenoptera, Apoidea: Megachilidae) of Fars province (Iran). Entomofauna 34: 229–260.

Kitamura, T. and Y. Maeta. 1969. Studies on the pollination of apple by Osmia. III. Preliminary report on the homing ability of Osmia cornifrons (Radoszkowski) and O. pedicornis Cockerell. Food and Agricultural Organization of the United Nations.

Koeniger N., G. Koeniger, M. Gries, S. Tingek, and A. Kelitu. 1996. Reproductive isolation of Apis nuluensis Tingek, Koeniger, and Koeniger, 1996 by species-specific mating time. Apidologie 27: 353–359.

Koeniger N., G. Koeniger, and S. Tingek. 2010. Honey bees of Borneo: exploring the centre of Apis diversity. Natural History Publications (Borneo), Kota Kinabalu xix+ [i]. 262p.

Koeniger, N., G. Koeniger, and D. Smith. 2011. Phylogeny of the genus Apis. In: Hepburn & Radloff (eds.): Honeybees of Asia. Springer-Verlag, Berlin. Pp: 23–50.

Koeniger N., G. Koeniger, S. Tingek, and A. Kelitu. 1996. Interspecific rearing and acceptance of queens between Apis cerana Fabricius, 1793 and Apis koschevnikovi Buttel-Reepen, 1906. Apidologie 27 (5): 371–380.

Koetz, A.H. 2013. Ecology, behaviour, and control of Apis cerana with a focus on relevance to the Australian incursion. Insects 4: 558–592.

Krombein, K.V. 1950. The Aculeate Hymenoptera of Micronesia II, Colletidae, Halictidae, Megachilidae, and Apidae. Proceedings of the Hawaiian Entomological Society 14: 101–142.

Krombein, K.V. 1967. Trap nesting wasp and bees: life histories, nests, and associates. Washington, DC: Smithsonian Press.

Krombein, K.V. 1969. Life history notes on some Egyptian solitary wasps and bees and their associates (Hymenoptera: Aculeata). Smithsonian Contributions to Zoology 19: 1–18.

Krombein, K.V. and B.B. Norden. 2001. Notes on trap-nesting Sri Lankan wasps and bees (Hymenoptera: Vespidae, Pompilidae, Sphecidae, Colletidae, Megachilidae). Proceedings of the Entomological Society of Washington 103: 274–281.

Krunic, M., M. Pinzauti, A. Felicioli, and L. Stanizavljevic. 1995. Further observations on Osmia cornuta Latr. and O. [Osmia] rufa L. as alternative fruit pollinators, domestication and utilization. Food and Agricultural Organization of the United Nations 47: 59–66.

Kuang, B.Y. and Y.C. Li. 1985. The genus Apis in China. Chinese Beekeeping 76: 7–9 (in Chinese).

Kuhlmann, M. and K. Timmerman. 2009. Nest architecture and floral hosts of the South African monolectic solitary bee Othinosmia (Megaloheriades) schultzei (Hymenoptera: Megachilidae). Entomologia Generalis 32: 1–9.

Kuhlmann, M., F.W. Gess, F. Koch, and S.K. Gess. 2011. Southern African osmiine bees: taxonomic notes, two new species, a key to Wainia, and biological observations (Hymenoptera: Anthophila: Megachilidae). Zootaxa 3108: 1–24.

Kurtak, B.H. 1973. Aspects of the biology of the European bee Anthidium manicatum (Hymenoptera: Megachilidae) in New York State. MS Thesis. Cornell University.


Laport, R.G. and R.L. Minckley. 2012. Occupation of active Xylocopa virginica nests by the recently invasive Megachile sculpturalis in upstate New York. Journal of the Kansas Entomological Society 85: 384–386.

Laroca S. and G.H. Rosado-Neto. 1975. Notas bionomicas: Hypanthidioides arenaria (Hymenoptera, Apoidea). Revista brasileira de biologia.

Li, S., Y.P. Meng, J.T. Chang, J.H. Li, S.Y. He, and B.Y. Kuang. 1986. A comparative study of esterase isozymes in 6 species of Apis and 9 genera of Apoidea. Journal of Apicultural Research 25: 129–133.

Linsley, E.G. 1946. Insect pollinators of alfalfa in California. Journal of Economic Entomology 39: 18–29.

Litman, J.R. 2012. Phylogenetic systematics and the evolution of nesting behavior, host-plant preference, and cleptoparasitism in the bee family Megachilidae (Hymenoptera, Apoidea). Cornell University 2012: 1–213.

Litman, J.R., T.L. Griswold, and B.N. Danforth. 2016. Phylogenetic systematics and a revised generic classification of anthidiine bees (Hymenoptera: Megachilidae). Molecular Phylogenetics and Evolution 100: 183–198. 

Lo, N., R. Gloag, D. Anderson, and B. Oldroyd. 2010. A molecular phylogeny of the genus Apis suggests that the giant honey bee of the Philippines, A. brevilingua Maa, and the plains honey bee of southern India, A. indica Fabricius, are valid species. Systematic Entomology 35: 226–233.

Lovell, J.H. 1909. The bees of Massachusetts: Osmia and Sphecodes. Entomological News 20: 122–126.


MacSwain, J.W. 1946. The nesting habits of Heteranthidium larreae. Pan-Pacific Entomologist 22: 159–160.

Maeta, Y. 1978. Comparative studies on the biology of the bees of the genus Osmia of Japan, with special reference to their managements for pollinations of crops (Hymenoptera: Megachilidae). Bulletin of the Tohoku National Agricultural Experiment Station. 57: 195–209.

Maier, C.T. 2009. New distributional records of three species of Megachilidae (Hymenoptera) from Connecticut and nearby states. Proceedings of the Entomological Society of Washington 111: 775–784.

Martin, D.J. 2009. Pollination and facultative ant-association in the African leopard orchid Ansellia africana. Journal of East African Natural History 98: 67–77.

Matsumoto, S. and T. Maejima. 2010. Several new aspects of the foraging behavior of Osmia cornifrons in an apple orchard. Psyche: 1–6.

McGinley, R.J. and J.G. Rozen Jr. 1987. Nesting biology, immature stages, and phylogenetic placement of the Palaearctic bee Pararhophites (Hymenoptera: Apoidea). American Museum Novitates 2903: 1–21.

Mello, M.L.S. and C.A. Garófalo. 1986. Structural dimorphism in the cocoon of a solitary bee, Lithurgus corumbae (Hymenoptera, Megachilidae) and its adaptive significance. Zoologischer Anzeiger 217: 195–206.

Michener, C.D. 1939. A revision of the genus Ashmeadiella (Hymen., Megachilidae). American Midland Naturalist 22: 1–84.

Michener, C.D. 1941. A synopsis of the genus Trachusa with notes on the nesting habits of T. perdita. Pan-Pacific Entomologist 17: 119–125.

Michener, C.D. 1944. The distribution of the osmiine bees of the deserts of North America. The American Naturalist 78: 257–266.

Michener, C.D. 1948. The generic classification of the anthidiine bees (Hymenoptera: Megachilidae). American Museum of Natural History, New York.

Michener, C.D. 1955. Some biological observations on Hoplitis pilosifrons and Stelis lateralis (Hymenoptera, Megachilidae). Journal of the Kansas Entomological Society 283: 81–87.

Michener, C.D. 1968. Nests of some African megachilid bees, with description of a new Hoplitis (Hymenoptera: Apoidea). Journal of the Society of the Entomological Society of Southern Africa 31: 337–359.

Michener, C.D. 1996. The first South African dioxyine bee and a generic review of the tribe Dioxyini (Hymenoptera, Megachilidae). Contributions on Hymenoptera and Associated Insects Dedicated to Karl V. Krombein, Memoirs of the Entomological Society of Washington 17: 142–152.

Michener, C.D. 2002. Duckeanthidium, a genus new to Central America, with generic synonymy and a new species (Hymenoptera: Megachilidae). Journal of the Kansas Entomological Society 75: 233–240.

Michener, C.D. 2007. The Bees of the World (2nd ed.). Johns Hopkins University Press, Baltimore and London, 953 pp.

Michener, C.D. and T.L. Griswold. 1994. The classification of old world Anthidiini. University of Kansas Science Bulletin 55: 299–327.

Michener, C.D. and E. Ordway. 1963. Some anthidiine bees from Mexico (Hymenoptera: Megachilidae). New York Entomological Society 72: 70–78.

Miguel I., M. Baylac, M. Oriondo, C. Manzano, L. Garnery, and A. Estonba. 2011. Both the geometric morphometric and microsatellite data consistently support the differentiation of the Apis mellifera evolutionary branch. Apidologie 42: 150–161.

Miller, S.R., R. Gaebel, R.J. Mitchell, and M. Arduser. 2002. Occurrence of two species of old world bees, Anthidium manicatum and A. oblongatum (Apoidea: Megachilidae), in Northern Ohio and Southern Michigan. The Michigan Entomology Society 35: 65–69.

Mitchell, T.B. 1933. A revision of the genus Megachile in the Nearctic region Part I: classification and descriptions of new species (Hymenoptera: Megachilidae). Transactions of the American Entomological Society 59: 295–361.

Mitchell, T.B. 1962. Bees of the eastern United States, 2ed. North Carolina Agricultural Experiment Station Technical Bulletin 152.

Mogga, J. and F. Ruttner. 1988. Apis florea in Africa: source of the founder population. Bee World 69: 100–103.

Morato, E.F. 2001. Biologia e ecologia de Anthodioctes moratoi Urban (Hymenoptera, Megachilidae, Anthidiini) em matas contínuas e fragmentos na Amazônia Central, Brazil. Revista Brasileira de Zoologia 18: 729–736.

Morawitz, F. 1876. Zur Bienenfauna der Caucasus Länder. Horae Societatis Entomologicae Rossicae (St. Petersburg) 12: 3–69.

Moure, J.S., and C.D. Michener. 1955. The bee family Fideliidae in South America (Hymenopt.-Apoidea). Dusenia 6: 199-206.

Moure, J.S. and D. Urban. 1994. Rhynostelis, gen. n. and taxonomic notes on Hoplostelis Dominique (Hymenoptera, Apoidea, Megachilidae). Revista Brasileira de Zoologia 11: 297–302.

Müller, A. 1996. Host-plant specialization in Western Palearctic anthidiine bees (Hymenoptera: Apoidea: Megachilidae). Ecological Monographs 66: 235–257.

Müller, A. 2006. Unusual host plant of Hoplitis pici, a bee with hooked bristles on its mouthparts (Hymenoptera: Megachilidae: Osmiini). European Journal of Entomology 103: 497–500.

Müller, A. 2012. Osmia (Orientosmia) maxschwarzi sp. n., a new Palaearctic osmiine bee with extraordinarily long mouthparts (Hymenoptera, Apiformes, Megachilidae). Mitteilungen der Schweizerischen Entomologischen Gesellschaft Bulletin De La Société Entomologique Suisse 85: 27–35.

Müller, A. 2018a. Palaearctic Osmia bees of the subgenus Hoplosmia (Megachilidae, Osmiini): biology, taxonomy and key to species. Zootaxa 4415: 297–329.

Müller, A. 2018. Palaearctic Osmiine Bees, ETH Zürich, http://blogs.ethz.ch/osmiini

Müller, A. and T.L. Griswold. 2017. Osmiine bees of the genus Haetosmia (Megachilidae, Osmiini): biology, taxonomy, and key to species. Zootaxa 4358 (2): 361–364.

Müller, A. and V. Trunz. 2014. Palaearctic Osmiine bees of the genera Hofferia and Stenoheriades (Megachilidae, Osmiini): biology, taxonomy, and key to species. Zootaxa 3765: 175–186.

Müller, A., A. Krebs, and F. Amiet. 1997. Bienen: Mitteleuropäische Gattungen, Lebensweise, Beobachtung. Naturbuch Verlag, Augsburg, pp. 384.

Müller, A., W. Topfl, and F. Amiet. 1996. Collection of extrafloral trichome secretions for nest wool impregnation in the solitary bee Anthidium manicatum. Naturwissenschaften 83: 230–232.

Murao, R., O. Tadauchi, and R. Miyanaga. 2015. The bee tribe Anthidiini (Hymenoptera, Megachilidae) collected from Central Asia. Japanese Journal of Systematic Entomology 21: 7–12.


Nadimi, A., A.A. Talebi, and Y. Fathipour. 2013. The tribe Osmiini (Hymenoptera: Megachilidae) in the north of Iran: new records and distributional data. Entomofauna 34: 205–220.

Nanork, P., S. Deowanish, and S. Wongsiri. 2001. Mitochondrial DNA variability of dwarf honey bee [Apis florea Fabricius (1787)] in Thailand by using PCR-RFLP technique. Proceedings of the 7th International Conference in Tropical Apiculture. Climates, Chiang Mai. Pp. 341–345.

Neff, J.L. and B.B. Simpson. 1992. Nest biology of Osmia (Diceratosmia) subfasciata Cresson in central Texas (Hymenoptera: Megachilidae). Pan-Pacific Entomology 68: 15–26.

Normandin, E., N.J. Vereecken, C.M. Buddle, and V. Fournier. 2017. Taxonomic and functional trait diversity of wild bees in different urban settings. PeerJ: 1–35.

Nunamaker, R.A., W.T. Wilson, and R. Ahmad. 1984. Malate dehydrogenase and non-specific esterase isoenzymes of Apis florea, A. dorsata, and A. cerana as detected by isoelectric focusing. Journal of the Kansas Entomological Society 57: 591–595.

Игнатенко, E.B. 2012. Кормовые связи пчёл (Hymenoptera: Apiformes) с цветковыми растениями в амурской области. Амурский зоологический журнал 9: 69–75.


O’Brien, M.F., D.R. Swanson, and J. Monsma. 2012. Anthidium oblongatum (Apoidea: Megachilidae) confirmed as a Michigan resident, with notes on other Michigan Anthidium species. The Great Lakes Entomologist 45: 102–105.

Oldroyd B. P., and P. Nanork. 2009. Conservation of Asian honey bees. Apidologie 40: 296–312.

Onuferko, T.M., R. Kutby, and M.H. Richards. 2015. A list of bee species (Hymenoptera: Apoidea) recorded from three municipalities in the Niagara region of Ontario, including a new record of Lasioglossum furunculum Gibbs (Halictidae) in Canada. JESO 146: 3–22.

Ornosa, C., F.J. Ortiz-Sanchez, and F. Torres. 2008. Catalogo de los Megachilidae del Mediterraneo occidental (Hymenoptera, Apoidea). III. Anthidiini y Dioxyini. Grasellsia 64: 51–86.

Ortiz-Sanchez, F.J. 2011. Lista actualizada de lase species de abejas de Espana (Hymenoptera: Apoidea: Apiformes). Boletin de la Sociedad Entomologica Aragonesa 49: 265–281.

Otis, G.W. 1991. A review of the diversity of species within Apis, in: Smith D. R. (Ed.), Diversity in the genus Apis. Westview Press, Boulder, Colorado. pp.: 29–49.

Otis, G.W. 1996. Distributions of recently recognized species of honeybees (Hymenoptera: Apidae: Apis) in Asia. Journal of the Kansas Entomological Society 69: 311–333.

Otis, G.W, S. Hadisoesilo, C. Sugiyani, and A.M. Cooper. 2001. Using social bees as indicators of habitat quality in Sulawesi, Indonesia. In: Proceedings of the 7th International Conference on Tropical Bees, Chiang Mai. Pp: 313–317.

Ozbek, H. and M. Schwarz. 2016. Distribution data of the kleptoparasitic bees in Turkey: part 2. Genera Coelioxys and Radoszkowskiana (Hymenoptera: Megachilidae: Megachilinae) with their checklist. Linzer Biol. Beitr. 48: 1753–1771.


Paar J, B.P. Oldroyd, E. Huettinger, and G. Kastberger. 2004. Genetic structure of an Apis dorsata population: the significance of migration and colony aggregation. Journal of Heredity 95 (2): 119–26.

Packer, L., J. Litman, and C.J. Praz. 2017. Phylogenetic position of a remarkable new fideliine bee from northern Chile (Hymenoptera: Megachilidae). Systematic Entomology 42: 473–488.

Parizotto, D.R. and G.A.R. Melo. 2015. Nests of the bees of the anthidiine genus Ananthidium Urban (Hymenoptera, Apidae, Megachilinae). Journal of Hymenoptera Research 47: 115–122.

Parizotto, D.R. and D. Urban. 2010. A new species of Dichanthidium (Hymenoptera, Megachilinae) and first records of the genus in Brazil. Revista Brasileira de Entomologia 54: 419–422.

Parker, F.D. 1975. Nests of the mason bees Osmia tanneri Sandhouse and O. longula Cresson with a description of the female of O. tanneri. Pan-Pacific Entomologist 51: 179–183.

Parker, F.D. 1977. Nests of Anthocopa enceliae (Cockerell) and A. elongate (Michener). Pan-Pacific Entomologist 55: 47–52.

Parker, F.D. 1984. Nesting biology of Osmia (Trichinosmia) latisulcata Michener. Journal of the Kansas Entomological Society 57: 430–436.

Parker, F.D. 1986. Nesting, associates, and mortality of Osmia sanrafaelae Parker. Journal of the Kansas Entomological Society 59: 367–377.

Parker, F.D. 1987. Nests of Callanthidium from block traps (Hymenoptera: Megachilidae). The Pan-Pacific Entomologist 63: 125–129.

Parker, F.D., J.H. Cane, G.W. Frankie, and S.B. Vinson. 1987. Host records and nest entry by Dolichostelis, a kleptoparasitic anthidiine bee. Pan-Pacific Entomologist 63: 172–177.

Partap, U. 1999. Conservation of endangered Himalayan honeybee, Apis cerana for crop pollination. Asian Bee Journal 1 (1): 44–49.

Partap, U. And L.R. Verma. 1998. Asian bees and bee keeping: issues and initiatives, Asian bees and bee keeping progress of research and development. Proceeding of fourth Asian Apicultural Association International Conference, Kathmandu, 23–28 March. Pp. 3–14.

Pasteels, J.J. 1969. La systématique générique et subgénérique des Anthidiinae (Hymenoptera, Apoidea, Megachilidae de l'ancien monde. Société royale d'entomologie de Belgique 31: 3–148.

Pasteels, J.J. 1972. Révision des Anthidiinae (Hymenoptera Apoidea) de la région Indo-malaise. Bulletin et Annuales de la société Royale Belge d’Entomologie 108: 72–128.

Pasteels, J.J. 1977. Une revue comparative de l’éthologie des Anthidiinae nidificateurs de l’ancien monde. Annales de la Société Entomologique de France 13: 651–667.

Pasteels, J.J. 1980. Revision du genre Euaspis Gerstäcker (Hymenoptera: Apoidea: Megachilidae). Bulletin & Annales de la Société Royale Belge d'Entomologie 116: 73–89.

Pauly, A., R.W. Brooks, L.A. Nilsson, Y.A. Pesenko, C.D. Eardley, M. Terzo, T.L. Griswold, M. Schwarz, S. Patiny, J. Munzinger, and Y. Barbier. 2001. Hymenoptera Apoidea de Madagascar et des îles voisines. Annales du Musée royal de l'Afrique centrale 286: 1–406.

Payne, A., D.A. Schildroth, and P.T. Starks. 2011. Nest site selection in the European wool carder bee, Anthidium manicatum, with methods for an emerging model species. Apidologie 42: 181–191.

Pechuman, L.L. 1967. Observations on the behavior of the bee Anthidium manicatum (L). Journal of the New York Entomological Society 75: 68–73.

Peisl, P. 1999. Beobachtungen und Gedanken zum Verhalten von Bienen-Mannchen. Bembix 12: 21–25.

Peters, D.S. 1978. Systematik und Zoogeographie der west-paläarktischen Arten von Osmia Panzer, 1806 s. str., Monosmia Tkalcu, 1974 und Orientosmia n. subgen. (Insecta: Hymenoptera: Megachilidae). Senckenbergiana Biologica (Frankfurt) 58: 287–346.

Phillips, J.K. and E.C. Klostermeyer. 1978. Nesting behavior of Osmia lignaria propinqua Cresson (Hymenoptera: Megachilidae). Journal of the Kansas Entomological Society 51: 91–108.

Pitts-Singer, T.L. and J.H. Cane. 2011. The alfalfa leafcutting bee, Megachile rotundata: the world’s most intensively managed solitary bee. Annual Review of Entomology 56: 221–37.

Ponomareva, A.A. 1958. Nesting habits of certain species of higher bees (Hymenoptera, Apoidea) in southwestern Turkmenistan. Entomologičeskoe obozrenie 37: 616–629.

Popov, V.V. 1961. On the evolution of the bee genera Protosmia Ducke and Chelostomopsis Cockerell. Zoologicheskii Zhurnal 40: 359–371.

Popov, V.B. 1967. The bees (Hymenoptera: Apoidea) of Iran. Trudy Zoologicheskogo Instituta Akademii nauk SSSR (Leningrad) 43: 184–216.

Praz, C. J. 2017. Subgeneric classification and biology of the leafcutter and dauber bees (genus Megachile Latreille) of the western Palearctic (Hymenoptera, Apoidea, Megachilidae). Journal of Hymenoptera Research 55: 1–54.

Praz, C.J., A. Muller, and S. Dorn. 2008. Specialized bees fail to develop on non-host pollen: do plants chemically protect their pollen? Ecology 38: 795–804.

Proctor, M., P. Yeo, and A. Lack. 1996. The natural history of pollination. Timber Press, Portland.

Prosi, R., H. Wiesbauer, and A. Muller. 2016. Distribution, biology and habitat of the rare European osmiine bee species Osmia (Melanosmia) pilicornis (Hymenoptera, Megachilidae, Osmiini). Journal of Hymenoptera Research 52: 1–36.


Quest, M. 2009. Arbestand, okologie und habitatwahl von bienen ausgewahlter offenlebensraume im lazovski zapovednik (ferner osten russland). Entomofauna (ansfelden), supplement 15: 1–357.


Radchenko, V.G. 1978. A new type of nests without cells in Metallinella atrocaerulea. Entomologicheskoe Obozrenie 57: 515–517 [In Russian; English translation in Entomological Review 57: 353–355.

Radloff S.E., H.R. Hepburn, and S. Fuchs. 2005. The morphometric affinities of Apis cerana of the Hindu Kush and Himalayan regions of western Asia, Apidologie 36: 25–30.

Radloff S.E., C. Hepburn, H.R. Hepburn , S. Fuchs, S. Hadisoesilo, K. Tan, M.S. Engel, and V. Kuznetsov. 2010. Population structure and classification of Apis cerana. Apidologie 41 (6): 589–601.

Radloff, S.E., H.R. Hepburn, and M.S. Engel. 2011. The Asian species of Apis. In: Hepburn, H. R. & S. E. Radloff (eds.). 2011. Honeybees of Asia. Springer-Verlag, Berlin. Chapter 1. Pp. 1–22.

Raffiudin, R. and R.H. Crozier. 2007. Phylogenetic analysis of honey bee behavioral evolution. Molecular Phylogenetics and Evolution 43: 543–552.

Rattawannee A., C. Chanchao, and S. Wongsiri. 2007. Morphometric and genetic variation of small dwarf honeybees Apis andreniformis Smith, 1858 in Thailand. Insect Science 14: 451–460.

Rightmyer, M.G., T. Griswold, and S.G. Brady. 2013. Phylogeny and systematics of the bee genus Osmia (Hymenoptera: Megachilidae) with emphasis on North American Melanosmia: subgenera, synonymies, and nesting biology revisited. Systematic Entomology 38: 561-576.

Rightmyer, M.G., M. Deyrup, J.S. Ascher, and T. Griswold. 2011. Osmia species (Hymenoptera, Megachilidae) from the southeastern United States with modified facial fairs: taxonomy, host plants, and conservation status. Zookeys 148: 257-278.

Rinderer, T. E. 1988. The rediscovery of Apis koschevnikovi. American Bee Journal 128: 807.

Rinderer, T.E., B.P. Oldroyd, S. Wongsiri, H.A. Sylvester, L.I. De Guzman, J.A. Stelzer, and R.M. Riggio. 1995. A morphological comparison of the dwarf honey bees of south-eastern Thailand and Palawan, Philippines. Apidologie 26: 387–394.

Rinderer, T.E., S. Wongsiri, B.Y. Kuang, J.S. Liu, B.P. Oldroyd, H.A. Sylvester, and L.I. De Guzman. 1996. Comparative nest architecture of the dwarf honey bees. Journal of Apicultural Research 35: 19–26.

Rinderer, T.E., B.P. Oldroyd, S. Wongsiri, H.A. Sylvester, L.I. De Guzman, S. Potichot, W.S. Sheppard, and S.L. Buchmann. 1993. Time of drone flight in four honey bee species in southeastern Thailand. Journal of Apicultural Research 32: 27–33.

Roberts, R.B. 1978. The nesting biology, behavior, and immature stages of Lithurge chrysurus, an adventitious wood-boring bee in New Jersey (Hymenoptera: Megachilidae). Journal of the Kansas Entomological Society 51: 735–745.

Rocha-Filho, L.C. and L. Packer. 2016. Phylogeny of the cleptoparasitic Megachilini genera Coelioxys and Radoszkowskiana, with the description of six new subgenera in Coelioxys (Hymenoptera: Megachilidae). Zoological Journal of the Linnean Society (2016: to be updated). DOI: 10.1111/zoj.12484

Romankova, T.G. 1985. New species of the genus Osmia (Hymenoptera, Megachilidae) from the Far East. Research Institute of Biology and Biophysics 722: 591.

Romankova, T.G. 1988. A new far-eastern bee of the tribe Anthidiini. Vestnik Zoologii 4: 25–30.

Romankova, T. 2003. Ontario nest-building bees of the tribe Anthidiini (Hymenoptera, Megachilidae). Journal of the Entomological Society of Ontario 134: 85–89.

Roubik, D.W., S. Sakagami, and I. Kudo. 1985. Note on distribution and nesting of the Himalayan honey bee Apis laboriosa Smith (Hymenoptera: Apidae). Journal of the Kansas Entomological Society 58 (4): 746–749.

Rozen, J.G. 1966. Taxonomic descriptions of the immature stages of the parasitic bee, Stelis (Odontostelis) bilineolata (Spinola) (Hymenoptera: Apoidea: Megachilidae). Journal of the New York Entomological Society 74: 85–91.

Rozen, J.G. 1973. Life history and immature stages of the bee Neofidelia. American Museum Novitates 2519: 1–14.

Rozen Jr., J.G. 1973. Immature stages of lithurgine bees with descriptions of the Megachilidae and Fideliidae based on mature larvae (Hymenoptera, Apoidea). American Museum Novitates 2527: 1–14.

Rozen, J.G. 1977. The ethology and systematic relationships of Fideliine bees, including a description of the mature larva of Parafidelia (Hymenoptera, Apoidea). American Museum Novitates 2637: 1–15.

Rozen Jr., J.G., 1987. Nesting biology of the bee Ashmeadiella holtii and its cleptoparasite, a new species of Stelis (Apoidea: Megachilidae). American Museum Novitates 2900: 1–10.

Rozen Jr., J.G. 1990. Pupa of the bee Pararhophites orobinus (Hymenoptera: Apoidea: Megachilidae). Journal of the New York Entomological Society 98: 379–382.

Rozen, J.G. 2015. Nest and immatures of the South American anthidiine bee Notanthidium (Allanthidium) chilense (Urban) (Apoidea: Megachilidae). American Museum Novitates 3826: 1–12.

Rozen, J. and G. Eickwort. 1997. The entomological evidence. Journal of Forensic Sciences 42: 394–397.

Rozen Jr. J.G. and M.S. Favreau. 1967. Biological notes on Dioxys pomonae pomonae and on its host, Osmia nigrobarbata (Hymenoptera: Megachilidae). Journal of the New York Entomological Society 75: 197–203.

Rozen, J.G. and G.H. Hall. 2012. Nesting biology and immatures of the oligolectic bee Trachusa larreae (Apoidea: Megachilidae: Anthidiini). American Museum Novitates 3765: 1–24.

Rozen Jr., J.G. and H.G. Hall. 2014. Nest site selection and nesting behavior of the bee Lithurgopsis apicalis (Megachilidae: Lithurginae). American Museum Novitates 3796: 1–24.

Rozen Jr., J.G. and S.M. Kamel. 2007. Investigations on the biologies and immature stages of the cleptoparasitic bee genera Radoszkowskiana and Coelioxys and their Megachile hosts (Hymenoptera: Apoidea: Megachilidae: Megachilini). American Museum Novitates 3573: 1–43.

Rozen, J.G., Jr. and S.M. Kamel. 2008. Hospicidal behavior of the cleptoparasitic bee Coelioxys (Allocoelioxys) coturnix, including descriptions of its larval instars (Hymenoptera: Megachilidae). American Museum Novitates 3636: 1–15.

Rozen Jr., J.G. and H. Özbek. 2004. Immature Stages of the cleptoparasitic bee Dioxys cincta (Apoidea: Megachilidae: Megachilinae: Dioxyini). American Museum Novitates 3443: 1–12.

Rozen Jr., J.G. and H. Özbek. 2005. Egg deposition of the cleptoparasitic bee Dioxys cincta (Hymenoptera: Apoidea: Megachilidae). Journal of the Kansas Entomological Society 78: 221–226.

Rozen, J.G. and C.J. Praz. 2016. Mature larvae and nesting biologies of bees currently assigned to the Osmiini (Apoidea: Megachilidae). American Museum Novitates 3864: 1–46.

Rozen Jr., J.G. and E.S. Wyman. 2013. Larval development and nesting biology of the adventive wood-boring bee Lithurgus (L.) chrysurus Fonscolombe (Hymenoptera: Megachilidae: Lithurgini). American Museum Novitates 3774: 1–20.

Rozen Jr., J.G. and E.S. Wyman. 2014. Early nesting biology of the wood-boring adventive bee, Lithurgus chrysurus Fonscolombe (Apoidea: Megachilidae: Lithurginae). American Museum Novitates 3804: 1–12.

Rozen Jr., J.G., H. Özbek, J.S. Ascher, C. Sedivy, C. Praz, A. Monfared, and A. Muller. 2010. Nests, petal usage, floral preferences, and immatures of Osmia (Ozbekosmia) avosetta (Megachilidae: Megachilinae: Osmiini), including biological comparisons with other osmiine bees. American Museum Novitates 3680: 1–22.

Rozen Jr., J.G., G. Pisanty, V. Trunz, D. Bénon, A. Dorchin, and C.J. Praz. 2015. Nesting biology, flower preferences, and larval morphology of the little-known old world bee Ochreriades fasciatus (Apoidea: Megachilidae: Megachilinae). American Museum Novitates 3830: 1–18.

Russo, L. 2016. Positive and negative impacts of non-native bee species around the world. Insects 7: 69.

Rust, R.W. 1974. The systematics and biology of the genus Osmia, subgenera Osmia, Chalcosmia, and Cephalosmia. Wasmann Journal of Biology 32: 1–93.

Rust, R.W. 1986. Biology of Osmia (Osmia) ribifloris Cockerell (Hymenoptera: Megachilidae). Journal of the Kansas Entomological Society 59 (1): 89–94.

Rust, R.W. and S.L. Clement. 1972. The biology of Osmia glauca and Osmia nemoris. Journal of the Kansas Entomological Society 46: 548–562.

Rust, R.W., G. Camon, J.T. Grossa, and B.E. Vaissière. 2004. Nesting biology and foraging ecology of the wood-boring bee Lithurgus chrysurus (Hymenoptera: Megachilidae). Journal of the Kansas Entomological Society 77: 269–279.

Ruttner, F. 1975. A metatarsal clasping structure in the Apis drone, Entomology Germany 2: 22–29.

Ruttner F., L. Tassencourt, and J. Louveaux. 1978. Biometrical-statistical analysis of the geographic variability of Apis mellifera L. Apidologie 9: 363–381.

Ruttner, F. 1988. Biogeography and taxonomy of honeybees. Springer, Berlin- New York. 284 pp.

Ruttner F., D. Kauhausen, and N. Koeniger. 1989. Position of the red honey bee, Apis koschevnikovi (Buttel-Reepen 1906), within the genus Apis. Apidologie 20 (5): 395–404.


Sakagami, S.F., T. Matsumura, and K. Ito. 1980. Apis laboriosa in Himalaya, the little known world largest honeybee (Hymenoptera, Apidae). Insecta Matsumurana 19: 47–78.

Sandhouse, G.A. 1939. The North American bees of the genus Osmia (Hymenoptera: Apoidea). The Entomological Society of Washington, Washington, D.C.: 1–172.

Sarzetti, L.C., J.F. Genise, and M.V. Sánchez. 2012. Trichothurgus bolitophilus sp. n. (Hymenoptera, Megachilidae) a bee nesting in horse manure pads in Patagonia, Argentina. Journal of Hymenoptera Research 29: 1–14.

Schrottky, C. 1902. Ensaio sobre as abelhas solitarias do Brazil. Revista do Museo Paulista 5: 330–613.

Schwarz, H.F. 1928. Bees of the subfamily Anthidiinae, including some new species and varieties, and some new locality records. Journal of the New York Entomological Society 36: 369–419.

Scott, V.L., S.T. Kelley, and K. Strickler. 2000. Reproductive biology of two Coelioxys cleptoparasites in relation to their Megachile hosts (Hymenoptera: Megachilidae). Annals of the Entomological Society of America 93: 941–948.

Scott, V.L., J.S. Ascher, T. Griswold, and C.R. Nufio. 2011. The bees of Colorado. Natural History Inventory of Colorado, 23, pp.1–100.

Sedivy, C., C.J. Praz, A. Müller, A. Widmer, and S. Dorn. 2008. Patterns of host-plant choice in bees of the genus Chelostoma: the constraint hypothesis of host-range evolution in bees. Evolution. 62: 2487–2507.

Sedivy, C., S. Dorn, and A. Muller. 2013. Molecular phylogeny of the bee genus Hoplitis (Megachilidae: Osmiini) – how does nesting biology affect biogeography? Zoological Journal 167: 28–42.

Sheffield, C., S. Dumesh, and M. Cheryomina. 2011. Hylaeus puntatus (Hymenoptera: Colletidae), a bee species new to Canada, with notes on other non-native species. Journal of the Entomological Society of Ontario 142: 29–43.

Sheppard, W. S. and M. D. Weixner. 2003. Apis mellifera pomonella, a new honey bee subspecies from Central Asia. Apidologie 34: 367–375.

Smith, F. 1858. Catalogue of the hymenopterous insects collected at Sarawak, Borneo: Mount Ophir, Malacca; and Singapore, by A. R. Wallace. Proceedings of the Linnean Society of London 2: 42–130.

Smith, F. 1861. Descriptions of new species of hymenopterous insects collected by A. R. Wallace at Celebes. Proceedings of the Linnean Society of London. 5: 57–93.

Smith, D.R. 1991. Mitochondrial DNA and honey bee biogeography, in: Smith D.R. (Ed.), Diversity in the genus Apis, Westview Press, Boulder, Co. pp. 131–176.

Smith D.R., L. Villafuerte, G.W. Otis G., and M. Palmer. 2000. Biogeography of Apis cerana F. and A. nigrocincta: insights from mtDNA studies. Apidologie 31 (2): 265–279.

Smith-Pardo, A.H. 2003. A preliminary account of the bees of Colombia (Hymenoptera: Apoidea): present knowledge and future directions. Journal of the Kansas Entomological Society 76: 335–341.

Somerville, D. 2011. Asian bees. Primefact 1093. Department of Industry and Investment, State of New South Wales (NSW), Australia. Available at http://www.dpi.nsw.gov.au/__data/assets/pdf_file/0003/382161/Asian-bees.pdf

Snelling, R.R. 1983. The North American species of the bee genus Lithurge (Hymenoptera: Megachilidae). Contributions to Science: Serial Publication of the Natural History Museum of Los Angeles County 343: 1–11.

Snelling, R.R. 1986. The taxonomic status of two North American Lithurge (Hymenoptera: Megachilidae). Bulletin of the Southern California Academy of Sciences 851: 29–34.

Snelling, R.R. 1987 A revision of the bee genus Aztecanthidium (Hymenoptera: Megachilidae). Pan-Pacific Entomologist 63: 165–171.

Snelling, R.R., 2003. Bees of the Hawaiian Islands, exclusive of Hylaeus (Nesoprosopis) (Hymenoptera: Apoidea). Journal of the Kansas Entomological Society 76 (2): 342–356.

Soh, E.J.Y., Z.W.W. Soh, S.X. Chui, and J.S. Ascher. 2016. The bee tribe Anthidiini in Singapore (Anthophila: Megachilidae: Anthidiini) with notes on the regional fauna. Nature in Singapore 9: 49–62.

Spear, D.M., S. Silverman, and J.R.K. Forrest. 2016. Asteraceae pollen provisions protect Osmia mason bees (Hymenoptera: Megachilidae) from brood parasitism. The American Naturalist 187: 797–803.

Stange, L.A. 1983. Synopsis of the genus Epanthidium Moure with the description of a new species from northeastern Mexico (Hymenoptera: Megachilidae). Pan-Pacific Entomologist 59: 281–297.

Steiner, V.B. and K.E. Whitehead. 1991. Resin collection and the pollination of Dalechampia capensis (Euphorbiaceae) by Pachyanthidium cordatum (Hymenoptera: Megachilidae) in South Africa 54: 67–72.

Stockl, P. 2000. Synopsis der Megachilinae Nord- und Sudtirols (Osterreich, Italien) (Hymenoptera: Apidae). Berichte des naturwissenschaftlichen-medizinischen Verein Innsbruck 87: 273–306.

Stubbs, C.S., F.A. Drummond, and E.A. Osgood. 1994. Osmia ribifloris biedermannii and Megachile rotundata (Hymenoptera: Megachilidae) introduced into the lowbush blueberry agroecosystem in Maine. Journal of the Kansas Entomological Society 67: 173–185.

Strange, J.P., J.B. Koch, V.H. Gonzalez, L. Nemelka, and T. Griswold. 2011. Global invasion by Anthidium manicatum (Linnaeus) (Hymenoptera: Megachilidae): assessing potential distribution in North America and beyond. Biological Invasions 13: 2115.

Struck, M. 1994. Flowers and their insect visitors in the arid winter rainfall region of southern Africa: observations on permanent plots. Insect visitation behavior. Journal and Arid Environments 28: 51–74.

Sulistiano, A. 1990. Morphometric analysis of Indonesian honeybees. Thesis. University of Wales College, Cardiff. 85p.


Tahmasebi G., R. Ebadi, N. Tajabadi, M. Akhoundi, and S. Faraji. 2002. The effects of geographical and climatological conditions on the morphological variation and separation of Iranian small honeybee (Apis florea F.) populations. Journal of Science and Technology of Agriculture and Natural Resources 6: 169–176.

Takahashi, J-I. 2005. Taxonomy and phylogeny of the genus Apis. Honeybee Science 26 (4): 145–152 (In Japanese with English abstract).

Takahashi J-I., J. Nakamura, M. Sasaki, S. Tingek, and S.I. Akimoto. 2002. New haplotypes for the non-coding region of mitochondrial DNA in cavity-nesting honey bees Apis koschevnikovi and Apis nuluensis. Apidologie 33 (1): 25–31.

Tan K., S. Fuchs, N. Koeniger, and Z. Ruiguang. 2002. Morphological characterization of Apis cerana (Fabr.) in the Province Yunnan of China. Apidologie 34: 553–561.

Tan K.,, X. Zhang, S. He, and D.Y. Zhou. 2003. Morphometrics and biogeography of Apis cerana in Yunnan, Apiculture China 53, 4–6 (in Chinese).

Tanaka H., D.W. Roubik, M. Kato, F. Liew, and G. Gunsalam. 2001. Phylogenetic position of Apis nuluensis of northern Borneo and phylogeography of A. cerana as inferred from mitochondrial DNA sequences. Insectes Sociaux 48: 44–51.

Tepedino, V.J., T.R. Toler, B.A. Bradley, J.L. Hawk, and T.L. Griswold. 2007. Pollination biology of a disjunct population of the endangered sandhills endemic Penstemon haydenii S. Wats. (Scrophulariaceae) in Wyoming, USA. Plant Ecology, 193: 59–69.

Teppner, H. 1996. Blüten und Blütenbesucher bei Onosma (Boraginaceae–Lithospermeae). Feddes Repertorium 106 (5–8): 525–532.

Thapa, R., R. Shrestha, D.N. Manandhar, S. Bista, and B. Kafle. 2001. Beekeeping in Nepal. In: Proceedings of the 7th International Conference on Tropical Bees. Chiang Mai. Pp. 409–413.

Thiele, R. 2002. Nesting biology and seasonality of Duckeanthidium thielei Michener (Hymenoptera: Megachilidae), an oligolectic rainforest bee. Journal of the Kansas Entomological Society 75: 274–282.

Thiele, R. 2005. Phenology and nest site preferences of wood-nesting bees in a Neotropical lowland rainforest. Studies on Neotropical Fauna and Environment 40: 39–48.

Tingek S., M. Mardan, G. Rinderer, N. Koeniger, and G. Koeniger. 1988. Rediscovery of Apis vechti (Maa, 1953): the Sabah honey bee. Apidologie 19: 97–102.

Tingek S., G. Koeniger, and N. Koeniger. 1996. Description of a new cavity-nesting species of Apis (Apis nuluensis n.sp.) from Sabah, Borneo, with notes on its occurrence and reproductive biology. Senckenbergiana Biologica 76: 115–119.

Tkalcu, B. 1969. Beiträge zur Kenntnis der Fauna Afghanistans. Osmiini, Megachilidae, Apoidea, Hym. Acta Musei Moraviae 54: 327–346.

Tkalcu, B. 1971. Zur Identität zweier Osmia-Arten (Hymenoptera, Apoidea, Megachilidae). Acta Entomologica Bohemoslovaca (Praha) 68: 222–230.

Tkalcu, B. 1974. Ergebnisse der Albanien-expedition 1961 des ‘Deutschen Entomologischen Institutes’. 89. Hymenoptera: Apoidea V (Megachilidae). Beiträge zur Entomologie Berlin 24: 323–348.

Tonietto, R.K. and J.S. Ascher. 2009. Occurrence of the old world bee species Hylaeus hyalinatus, Anthidium manicatum, A. oblongatum, and Megachilidae sculpturalis, and the native species Coelioxys banksi, Lasioglossum michiganense, and L. zophops in Illinois (Hymenoptera: Apoidea: Colletidae, Halictidae, Megachilidae). The Great Lakes Entomologist 41: 200–203.

Torchio, P.F. 1976. Use of Osmia lignaria Say (Hymenoptera: Apoidea, Megachilidae) as a pollinator in an apple and prune orchard. Journal of the Kansas Entomological Society 49: 475–482.

Torchio, P.F. 1990. Osmia ribifloris, a native bee species developed as a commercially managed pollinator of highbush blueberry (Hymenoptera: Megachilidae). Journal of the Kansas Entomological Society 63: 427–436.

Torchio, P. and E. Asensio. 1985. The introduction of the European bee, Osmia cornuta Latr, into the US as a potential pollinator of orchard crops, and a comparison of its manageability with Osmia. Journal of the Kansas Entomological Society 58: 42–52.

Trung, L.Q., P.X. Dung, and T.X. Ngan. 1996. A scientific note on first report of Apis laboriosa F Smith, 1871 in Vietnam. Apidologie 27: 487–488.

Trunz, V., L. Packer, J. Vieu, N. Arrigo, and C.J. Praz. 2016. Comprehensive phylogeny, biogeography and new classification of the diverse bee tribe Megachilini: Can we use DNA barcodes in phylogenies of large genera? Molecular Phylogenetics and Evolution 103: 245–259.

Turell, M.J. 1976. Observations on the mating behavior of Anthidiellum notatum and Anthidiellum perplexum. The Florida Entomologist 59: 55–61.


Underwood, B. A. 1986. The natural history of Apis laboriosa Smith in Nepal. M.Sc. Thesis, Cornell University, Ithaca, NY. 88p.

Underwood, B.A. 1990. Time of drone flight in Apis laboriosa Smith in Nepal. Apidologie 21: 501–504.

Ungricht, S., Müller, A. and Dorn, S. 2008. A taxonomic catalogue of the Palaearctic bees of the tribe Osmiini (Hymenoptera: Apoidea: Megachilidae). Zootaxa 1865: 1–253.

Urban, D. 1992. Considerações sobre Anthidulum Michener, stat. n. e Dicranthidium Moure & Urban, stat.n. e descrições de espécies novas (Hymenoptera, Megachilidae). Revista Brasileira de Zoologia 9: 11–28.

Urban, D. 1997. Larocanthidium gen. n. de Anthidiinae do Brasil (Hymenoptera, Megachilidae). Revista Brasileira de Zoologia 14: 299–317.

Urban, D. 2002. Duas espécies novas de Dicranthidium Moure & Urban (Hymenoptera, Megachilidae) e chave para a identificação das espécies. Revista Brasileira de Zoologia 19: 637–643.

Urban, D. 2004. Espécies novas e notas sobre Dianthidiini (Hymenoptera, Megachilidae) do Peru. Revista Brasileira de Entomologia 48: 347–352.

Urban, D. and J. Moure. 2012. Anthidiini Ashmead, 1899. In Moure, J. S., Urban, D., and Melo, G. A. R. (Orgs.). Catalogue of Bees (Hymenoptera, Apoidea) in the Neotropical Region. 

Urban, D. and D.R. Parizotto. 2011. Michanthidium almeidai, a new species from northeastern Brazil (Hymenoptera: Megachilidae. ZooKeys 132: 65–73.

Urban, D. and D.R. Parizotto. 2012. A revised key to the Neotropical cleptoparasitic anthidiine genera (Hymenoptera, Megachilinae) with notes and descriptions of the male of Rhynostelis Moure & Urban. Zookeys 249: 27–35.


Vicens, N., Bosch, J. and Blas, M. 1994. Biology and population structure of Osmia tricornis Latreille (Hym., Megachilidae). Journal of Applied Entomology 117: 300–306.

Villalobos E.M. and T.E. Shelly. 1991. Correlates of male mating success in two species of Anthidium bees (Hymenoptera: Megachilidae). Behavioral Ecology and Sociobiology 29: 47–53.


Wainwright, C.M. 1978. Hymenopteran territoriality and its influence on the pollination ecology of Lupinus arizonicus. The Southwestern Naturalist 23: 605–616.

Warncke, K. 1980. Die Bienengattung Anthidium Fabricius, 1804 in der Westpalaarktis und im turkestanischen Becken. Entomofauna 1: 119–210.

Wei, S.G., Wang, R., Smirle, M.J., Xu, H.L. 2002. Release of Osmia excavata and Osmia jacoti (Hymenoptera: Megachilidae) for apple pollination. Canadian Entomologist 134: 369–380.

Westrich, P. 1989. Die Wildbienen Baden-Württembergs. Eugen Ulmer GmbH & Co., Stuttgart

Westrich, P. 1990. Die Wildbienen Baden-Wurttembergs. Spezieller Teil: Die Gattungen und Arten. Verlag Eugen Ulmer. Pp. 437–972.

Whitehead, V.B. 1984. Distribution, biology, and flower relationships of fideliid bees of southern Africa (Hymenoptera, Apoidea, Fideliidae). African Zoology 19: 87–90.

Wilson, J.S., L.E. Wilson, L.D. Loftis, and T. Griswold. 2010. The montane bee fauna of north central Washington, USA, with floral associations. Western North American Naturalist, 70: 198–207.

Wilson, J.S. and O.M. Carril. 2016. The Bees in Your Backyard: A Guide to North America’s Bees. Princeton University Press, Princeton and Oxford, 288 pp.

Winston, M. L. 1991. The biology of the honey bee. Harvard University Press. 249p.

Wirtz, P., H. Szabados, H. Pethig, and J. Plant. 1988. An extreme case of interspecific territoriality: male Anthidium manicatum (Hymenoptera: Megachilidae) wound and kill intruders. Ethology 78: 159–167.

Wongsiri, S., K. Limbipichai, P. Tangkanasing, M. Mardan, T. Rinderer, H.A. Sylvester, G. Koeniger, and G.W. Otis. 1990. Evidence of reproductive isolation confirms that Apis andreniformis (Smith, 1858) is a separate species from sympatric Apis florea (Fabricius, 1787). Apidologie 21: 47–52.

Wongsiri, S.P, C. Lekprayoon, R. Thapa, K. Thirakupt, T.E. Rinderer, H.A. Sylvester, B.P. Oldroyd, and U. Boomcham. 1996. Comparative biology of Apis andreniformis and Apis florea in Thailand. Bee World 77 (4): 23–35.

Wu, Y. 2004. Ten new species of the tribe Osmiini from China (Apoidea, Megachilidae, Osmiini). Acta Zootaxonomica Sinica 29: 531–537.

Wu, Y. 2006. Hymenoptera: Megachilidae. Fauna Sinica, Insecta. Vol. 4. Science Press, Beijing.

Wu, Y.R. and B. Kuang. 1986. A study of the genus Micrapis (Apidae). Zoological Research 7: 99–102 (in Chinese).

Wu, Y.R. and B. Kuang. 1987. Two species of small honeybee – a study of the genus Micrapis. Bee World 68: 153–155.


Yasumatsu, K. and Hirashima, Y. 1950. Revision of the genus Osmia of Japan and Korea (Hymenoptera: Megachilidae). Mushi (Fukuoka) 21: 1–18.

Yanega, D. 1994. Nests and hosts of three species of megachild bees (Hymenoptera: Apoidea: Megachilidae) from Coahuila, México. Journal of the Kansas Entomological Society 67: 415–417.


Zanella, F.C.V. and A.G. Ferreira. 2005. Registro de hospedeiro de Austrostelis Michener & Griswold (Hymenoptera: Megachilidae) e de sua ocorrência na Caatinga. Neotropical Entomology 34: 857–858.

Zhen Ming J., Y. Guanhuang, H. Shuanghiu, L. Shikui, and R. Zaizin. 1992. The advancement of apicultural science and technology in China. In: Verna L. R. (ed.): Honey bees in mountain agriculture. Oxford and IBH Publishing, New Delhi. Pp. 133–148.